THE VACUOME THEORY 



101 



Gatenby (30b) shows that it is undoubtedly pecuHar in many 

 features. The Golgi apparatus is apparently represented by the 

 so-called " Nebenkern." This consists of a spherical mass of 

 differentiated cytoplasm surrounded by, or rather containing at 

 its periphery, a number of batonettes (dictyosomes). These 

 obviously correspond to the Golgi bodies of other forms, the central 

 mass being idioplasm. The situation is complicated by the fact 

 that two kinds of mitochondria occur, the micromitosomes and 



A B c 



Fig. 53. — Diagrams of mucous cells from the gastric epithelium of 

 Triton to illustrate the relation of the vacuome to the classical 

 Golgi apparatus (from Parat and Painleve (109) ). A, by 

 Dietrich's method ; " chondriocontes," black ; vacuome, white. 

 B, by Da Fano's method ; the classical Golgi apparatus. C, 

 after intra vitani staining in neutral red ; the vacuome. 



the macromitosomes. These two sets take part in the formation 

 of the tail sheaths, the former making the front sheath, the latter 

 the hind sheath. The " Nebenkern " does not take any part in 

 the formation of the tail sheath. It appears to be sloughed off. 

 The formation of the acrosome is not observed. 



Parat denies that the dictyosomes are the real Golgi apparatus. 

 This latter is really represented, he says, by the vacuoles which 

 lie in the archoplasm (idioplasm), which stain in neutral red. 

 The batonettes or cortex of the archoplasm are really the " lepido- 

 some " of mitochondria. This criticism is, of course, based on the 



