GOLGI IN PLANTS 



103 



claiming any definite homology for them, maintains that they 

 introduce a new complexity into the problem. No longer is it a 

 case of homologising two structures in both animals and plants. 

 These platelets do not stain either in Janus green or in neutral red, 

 and, therefore, whatever they may be, they belong to neither 

 of the two previously recognised systems : the chondriome 

 and vacuome. Later he has amplified the description, and 

 Gatenby and others (110) have confirmed this very fully. The 

 latter are quite definite in that they consider that the plant 

 osmiophilic platelets are Golgi apparatus as they are demonstrated 

 by Kolatchev and Mann-Kopsch methods. So far Benda, 



■?.N--.V. o;,^V ■:.,■. 0*. . o ••.'r/--..-.:oo-- 



%--'iW^:c6'' 



.'.^■■'■\o- ■ V t 



i-.- .■"■■>•:>«■{> *>'*v«''' / 



Fig. 54, — Meristem cells from root tips after Kolatchev's method 

 (from Bowen (12) ). A, in Barley : B, in Kidney bean ; P., osmio- 

 philic platelets ; A\, nucleus ; l^ac, vacuoles. 



Flemming- without-acetic and hematoxylin, Champy-hsematoxylin 

 and silver-nitrate methods have failed, but few modifications have 

 been tried as yet. 



Gatenby does not agree with Bowen as to the exact interpreta- 

 tion of the vacuome in plants. He considers that Bowen's 

 vacuome is probably due to corrosive-osmium artefacts, and that 

 the true vacuoles may be associated with the Golgi apparatus, 

 although they are certainly not " protoplasmic inclusions," but 

 " deutoplasmic." 



This discovery, then, deals a very decided blow at the vacuome 

 theory. The final disproof has been forthcoming quite recently in 

 a paper by Gatenby (35), in 1929, in which he has studied the 



