Francis E. Lloyd —96— Carnivorous Plants 



in young organs lie at the same level. With maturity, the epidermis 

 becomes ribbed with sunken furrows between the ribs. The floor of 

 the furrow is composed of a double row of shorter cells, each pair 

 bearing a sessile gland (74 — 10, 13). In scattered positions occur 

 stalked glands which secrete abundant mucilage. In the sepals the 

 epidermal cells are wavy-walled on both surfaces, less so on the outer 

 (lower) surface toward the base. On the outer, dorsal face of the leaf 

 occur both sessile and stalked glands, the latter very numerous, on the 

 inner face only sessile glands occur (p — 9). Stalked glands are to be 

 found even on the ovary wall. Stomata occur on both faces of the 

 sepals, and on the leaves and stem they are to be found interrupting 

 the rows of sessile glands {14 — 13). They are somewhat raised and 

 extend considerably above the ditch bottom. In this way according to 

 Fenner the stomatal pore does not become clogged with the secretion 

 of the sessile glands, which probably fills that reach of the ditch oc- 

 cupied by them. 



The leaves are long, slender and linear in form, tapering toward 

 the apex. When in the bud they display, in the case of B. gigantea, 

 no circination, the apices showing only a very meagre outward curva- 

 ture, if any. In B. Hnifolia, however, the leaves are outwardly cir- 

 cinate, as in Drosophyllum. This somewhat surprising fact was clearly 

 seen in the material from Arnhem Land sent me by my friend Mr. 

 Charles Barrett, and figured in 14 —7. B. gigantea is seen in 14 — 

 8. Of this DiELS (1930) says merely that the leaves are spirally in- 

 rolled at the tip. 



In transverse section the leaves are triangular with round angles 

 {14 — 11). Toward the tip they become nearly cylindrical and the tip 

 itself is somewhat enlarged to form a knob, properly interpreted to be 

 a hydathode (Lang, Fenner). Its interior is occupied by a large 

 mass of tracheidal tissue in contact with and ending the vascular 

 strand which reaches thereto. One or two protuberant stomata are to 

 be found at the apical surface, not by any means always at the extreme 

 apex, together with both stalked and sessile glands. The rigidity of 

 the leaf, which is very slender for its length, is attained by the very 

 thick-walled epidermis and the strands of mechanical tissue accom- 

 panying the vascular bundles. Beneath the epidermis on all sides 

 there is a thick layer of chlorenchyma in which there is no sharp de- 

 markation between palisade and spongy tissue. All of the cells are 

 oval rather than columnar and lie in three courses. Beneath the 

 epidermis the palisade cells have expanding ends in contact with it 

 {14 — 9, 10). This, Fenner explains, ensures a contact for lively 

 diffusion between the glands and the vascular system. The upper leaf 

 surface is rather flat, with a very shallow depression along the middle. 

 On this surface there are very few stalked glands, which on the lower 

 surface are very numerous. Sessile glands are as numerous here as 

 elsewhere {g — 9) . 



The sessile gland {14 — 10, 13) stands upon a pair of epidermal 

 cells, and consists of a capital of eight radially disposed cells, supported 

 on a single very short stalk cell, this resting on two short epidermal 

 cells, which according to Fenner originate from a single basal cell of 

 the very young trichome. The furrow in which the sessile glands stand 



