l6 BOTANICAL GAZETTE [july 



at the angle of junction between pileus and stipe. This soon 

 becomes a definite level palisade layer. It is entirely exogenous 

 in origin. 



3 . There is a strong epinastic development in the margin of the 

 pileus and it becomes so strongly incurved that the filaments on its 

 margin intermingle with those on the surface of the stipe. This 

 occurs while the hymenophore is still in a level palisade condition. 



4. The gills originate as downward growing folds which 

 develop centrifugally, the first folding taking place at the point 

 where the fundament of the hymenophore was first distinguishable 

 over the angle of stipe and pileus. 



5. The secondary gills originate in the same manner as the 

 primary gills but at varying distances from the stipe. Their 

 development is centrifugal. 



6. The primary gills during their early development are attached 

 to the stem and only become free during the final expansion of the 

 fruit body. 



7. The cystidia are distinguishable as soon as the gill salients 

 appear. They appear as larger cells with scanty protoplasmic 

 content, while the smaller cells of the hymenial layer are densely 

 filled with protoplasm. 



8. The cystidia are formed terminally upon filaments similar 

 to, but usually larger than, those that bear the smaller cells of the 

 hymenial layer. In younger stages the filaments bearing cystidia 

 are little if at all branched, but in older fruit bodies they become 

 more branched. The filaments bearing basidia and paraphyses 

 branch profusely very early in their development. 



9. The surface of the pileus is covered with cells that are similar 

 to the cystidia. 



10. The trama in the young gills is composed of a few slender 

 filaments which lie more or less parallel to each other. During 

 expansion large elongated cells developing from the subhymenium 

 grow inward and downward, giving a very unusual appearance to 

 the trama. These cells probably represent internal cystidia. The 

 cells of the original trama become much enlarged also. 



11. The cells in all parts of the young basidiocarps are con- 

 stantly binucleate. 



