iqiq] HAYATA—PROTOMARATTIA 87 



A . Henryi is only known from Mentzu (Yunnan) ; A . Somai exists 

 in one or two spots in the northern part of Formosa ; A . subintegra 

 occurs in one place in the mountains of Chapa between Yunnan 

 and Tonkin; and A. tamdaoensis is found in one locality on Mt. 

 Tamdao (Tonkin) ; and the new genus is also limited to one spot 

 on Mt. Tamdao. 



Returning to the systematic position of the new genus, the most 

 remarkable feature which separates the new type from all the other 

 genera of the Marattiaceae is its elongated linear or even vermiform 

 synangium. The other important characters are its horizontal 

 dorsiventral rhizomes and simply pinnate fronds. Through the 

 dorsiventral rhizomes it is related to Kaulfussia and Archangiop- 

 teris; by the simple pinnate fronds it is aUied to Macroglossum, 

 Danaea, and Archangiopteris; in the structure of the synangium 

 its nearest of kin is to be found in Marattia. It is distinguishable 

 from the latter, however, by its elongated linear synangium, simply 

 pinnate fronds, and dorsiventral horizontal rhizome. It differs from 

 Archaiigio pteris in having a synangium; from Macroglossum in the 

 dorsiventral rhizomes and synangium; from Angiopteris in the 

 simple pinnate leaves, dorsiventral rhizomes, and synangium; from 

 Danaea in the synangium with a longitudinal common slit; and 

 from Kaulfussia in the pinnate leaves and linear synangium. After 

 considering all these cases, I am forced to the conclusion that the 

 new plant must be a type representing a new genus. I propose 

 to call it ProtomaraUia, as it bears exactly the same relation to 

 Marattia as Archangiopteris does to Angiopteris. 



As was stated, Protomarattia closely resembles Marattia in the 

 reproductive organs, while it is closely related to Archangiopteris 

 in its vegetative organs. The similarity of the type of the new 

 genus and Archangiopteris tamdaoensis in the fronds and rhizomes, 

 even in the serration and venation, is really so very great that I 

 entirely failed to distinguish the one from the other until I saw the 

 sori. The protective arrangement of stipules and commissures 

 of our plant is exactly like that in Archangiopteris. The synangium 

 also, presenting a linear form, with comparatively thinner lateral 

 walls and a little looser connection of locules, more or less tends 

 toward the sorus of Archangiopteris, or even toward that of 



