PAPENFUSS: CLASSIFICATION OF THE ALGAE 187 



Phylogeneticall}^ the red algae appear to be distantly related to the blue- 

 green algae by way of the Bangiophycidae. Cohn (1867) was the first to arrive 

 at this conclusion on the basis of his investigation of the pigments of both 

 groups. Such a relationship appeared likely to Berthold (1882) also and is ac- 

 cepted by Ishikawa (1921, 1924), Kylin (1930b, 1937a, 1943b), Tilden (1933, 

 1935) and Skuja (1938). 



In the synoptic outline that follows, Skuja's (1939a) classification of the 

 Bangiophycidae (which has been accepted by Kylin, 1944a, and Tanaka, 1952) 

 has been followed. The classification of the Florideophycidae is essentially that 

 of Kylin (1932). 



Phylum RHODOPHYCOPHYTA Papenfuss (1946, p. 218) 

 Syn.: Rhodophyta Wettstein (1901, p. 46) 

 Class RiionopiiYCEAE Ruprecht (1851, p. 205) 



Syn.: Rhodospei-meae Harvey (1836, p. 160) ; Heterocarpeae Kiitzing (1843, p. 369) ; 

 Pliycoerytlii'inophycees Marchand (1895, p. 17) 



Subclass BANGIOPHYCIDAE De Toni orth. mut. L. M. Newton (1953, p. 406) 



Syn.: Bangioideae De Toni (1897, p. 4) ; Protoflorideae Rosenvinge (1909, p. 55) 

 Order PORPHYRIDIALES Kylin (1937b, p. 4; see also Kylin, 1937a, pp. 39- 

 51) 



Family Porphyridiaceae Kylin (1937b, p. 4) 

 Order GONIOTRICHALES Skuja (1939a, p. 31) 



Family Goniotrichaceae (Rosenvinge) Skuja (1939a, p. 31) 

 Family Phragmonemataceae Skuja (1939a, p. 32) 

 Order BANGIALES Engler (1892, p. 15) 



Family Erythropeltidaceae Skuja (1939a, p. 33) 

 Family Bangiaceae (S. F. Gray) Nageli (1847, p. 136) 



Syn.: Porphyraceae Kiitzing orth. mut. Rabenhorst (1868, p. 397); 

 Erythrotrichiaceae Marchand orth. mut. G. M. Smith (1944, p. 162) 

 Order RHODOCHAETALES Skuja (1939a, p. 34) 



Family Rhodocliaetaceae Schmitz, in Schmitz and Hauptfleisch (1896, 

 p. 317) 

 Order COMPSOPOGONALES Skuja (1939a, p. 34) 



Family Compsopogonaceae Schmitz, in Schmitz and Hauptfleisch (1896, 

 p. 318) 

 Subclass FLORIDEOPHYCIDAE (Lamouroux) Engler orth. mut. L. M. Newton (1953, 

 p. 407) 



Syn.: Floridees Lamouroux (1813, p. 115); Euflorideae Johnson (1894, p. 639) 

 Order NEMALIONALES Schmitz, in Engler (1892, p. 17) 

 Syn.: Bonnemaisoniales Feldmann (1952, p. 29) 

 Family Acrochaetiaceae Fritsch (1944, p. 258) 



Syn.: Chantransiaceae Kiitzing orth. mut. Rabenhorst (1868, p. 400; 

 not including Chantransia De Candolle, see Silva, 1952, pp. 261-262); 

 Rhodochortonaceae Nasr (1947. p. 92) 

 Family Batrachospermaceae (C. Agardh) Dumortier orth. mut. Raben- 

 horst (1868, p. 404) 



Family Lemaneaceae (S. F. Gray) Harvey orth. mut. Rabenhorst (1863, 

 p. 275) 



Family Helminthocladiaceae (J. Agardh) Harvey orth. mut. Hauck 

 (1883, p. 14) 



Syn.: Nemalionaceae Cohn orth. mut. G. Murray (1895, p. 207) 

 Family Chaetangiaceae Kiitzing orth. mut. Hauck (1883, p. 14) 

 Family Thoreaceae Reichenbach ex Hassall orth. mut. Schmitz, in 

 Schmitz and Hauptfleisch (1896, p. 321) 

 Family Naccariaceae Kylin (1928, p. 11) 

 Family Bonnemaisoniaceae Schmitz, in Engler (1892, p. 20) 



