38 



OCEANIC TINTINNOINA OF LAST CRUISE OF CARNEGIE 



Dictyocysta tiara Hacckel 

 Dictyocysta tiara, Kofoid and Campbell, 1929, p. 302, fig. 567. 



The rather moderately tall, narrow lorica, with single row 

 of fenestrae and elongated bowl, has a length of 2.0 oral 

 diameters. The oral margin is undulating, with 8 very flat 

 arches. The collar is cylindrical, 0.75 oral diameter in length, 

 and has 8 rectangular, subequal open windows with rounded 

 lower and angular upper corners. The bowl has a narrow 

 shoulder, reaches a diameter of 1.25 oral diameters at the 

 shoulder, which is a little lower than the lower end of the 

 collar, then contracts (30 ) for two-thirds of its length, and 

 finally becomes subcorneal (98 ). The aboral end is rounded. 



The wall of the bowl has a single row of 8 elongated, 

 narrow oval fenestrae in the equatorial region. More 

 numerous tiny fenestrae occur in a double row below the 

 major row. The remaining surface of the bowl and the 

 supporting beams and mullions of the collar are studded 

 with closely set, minute coccoliths. The collar windows are 

 open. 



Length, 8o|i. 



This much misunderstood species is more elongated and 

 narrower than any other. 



Recorded from four stations in the Pacific, as follows: one 

 (45) in the Galapagos region, one (64) in the South Pacific 

 middle latitudes, and two (in, 112) in the North Pacific 

 middle latitudes. 



There are 5 pump samples, of which 1 was taken at the 

 surface and 2 each at 50 and 100 meters. Frequency, 2 per 

 cent at stations 45, 64, m; average, 1.6 per cent. 



Temperature, i5?oo-22?43 ( ig?gi ); salinity, 34.54-35.26 

 (34.73); density, 24.02-25.43 (24.58); pH, 8.10-8.22 (8.16). 



COXLIELLIDAE Kofoid and Campbell emended 

 Coxliellidae, Kofoid and Campbell, 1929, p. 91. 



The family includes four genera: Coxliella, Climacocylis, 

 Helicostomella, and Metacylis. Only the last does not occur 

 in the material of this expedition. All but Helicostomella 

 are eupelagic, the latter being coastal; most of them are 

 tropical. 



Coxliellinae Kofoid and Campbell 

 Coxliellinae Kofoid and Campbell, 1939, p. 90. 



The Coxliellinae include those members of the family in 

 which the spiral lamina extends the whole length of the 

 lorica, except in some species in which it may be limited. 



The genera included are Coxliella and Climacocylis. 



COXLIELLA Brandt emended 

 Coxliella, Kofoid and Campbell, 1929, pp. 95-96. 



Coxliella has a spiral lamina which forms the bowl and 

 horn. This spiraling is also found in the collar of Codonel- 

 lopsis and in a few species of Tintinnopsis. It is not un- 

 likely that Coxliella had its origin in the latter genus, and 

 that the spiral collar of Codonellopsis is of a different source. 

 The wall is rigid, the flaccidity of that of Climacocylis being 

 derived. The aboral end of Coxliella is closed, that of 



Climacocylis open; this is an additional suggestion of the 

 derivation of the latter genus. 



Species of Coxliella are common in warm and temperate 

 seas. Only the subgenus Protocochliella is from cool waters 

 in both hemispheres, and is not found in the tropical ocean. 



Six species are described here. 



Coxliella declivis Kofoid and Campbell 

 Coxliella declivis Kofoid and Campbell, 1929, p. 97, fig. 192. 



The tapering subconical lorica, with few turns and narrow 

 aboral horn, has a length of 1.89 to 1.93 oral diameters. The 

 oral margin is entire, thin, and smooth. The short, stout 

 bowl is cylindrical in the upper fourth, and convex conical 

 in the lower three -fourths; the upper section is 55 to 6o°, 

 whereas near the horn it is 95 . The short, conical aboral 

 horn is somewhat under 0.3 oral diameter in length; it is 

 twisted, and its end is minutely blunted. 



The wall reaches as much as 0.09 oral diameter in thick- 

 ness across the widest of the 9 to 10 left-turning spiral whorls. 

 The fifth and sixth turns reach a width of 0.33 oral diameter; 

 the uppermost and lowermost ones are only half that much, 

 or less. There are thin laminae with three to five layers of 

 tiny alveoles closely packed within their confines. The wall 

 is dull, and lacunae are absent. 



Length, nou,. 



Coxliella declivis is more tapering and has a more sym- 

 metrical aboral horn than C. laciniosa. Its wide turns sug- 

 gest C. pseudannulata, but its horn is decidedly different, 

 and neither is it like that of C. pelagica. Coxliella decipiens 

 is longer, with more and narrower spiral turns and some- 

 what similar aboral end. The remaining species are all so 

 different that no difficulty in identification should arise. 



Recorded from seven stations, two in the Atlantic and 

 five in the Pacific, as follows: two (18, 19) in the Sargasso 

 Sea, two (41, 85) in the Galapagos region, one (99) in the 

 Pacific equatorial region, and two (133, 134) in the Cali- 

 fornia region. 



There are 2 pump and 6 net samples, of which 2 were 

 taken at the surface, 2 at 50 meters, and 4 at 100 meters. 

 Maximum frequency, 3 per cent at station 99; other records 

 at 2 per cent from stations 19, 85. 



Temperature: Atlantic, net samples 20?32-25?3i (22?68); 

 Pacific, pump samples i8?i8-22?68 (20^43), net samples 

 20?42-27?89 (25^38). Salinity: Atlantic, net samples 36.81- 

 37.15 (37.00); Pacific, pump samples 34.63-34.70 (34.66), 

 net samples 34.19-36.24 (35.15). Density: Atlantic, net 

 samples 24.80-26.07 (25.48); Pacific, pump samples 23.82- 

 24.96 (24.39), net samples 22.50-24.06 (23.31). pH: At- 

 lantic, net samples 8.21-8.27 (8-24); Pacific, pump samples 

 8.34-8.47 (8.40), net samples 8.1 1-8.22 (8.18). 



Coxliella fasciata (Kofoid) Brandt 

 (Figures 30, 34) 



Coxliella fasciata, Kofoid and Campbell, 1929, pp. 97-98, fig. 



205. 



The greatly elongated conical lorica, with few spiral turns 

 and slight flare, and with narrowed aboral end, has a length 



