Cytology and Movements of the Cyanophycece. 255 



mal structural appearance in the Cyanophycean cell. They 

 are generally to be seen in all Cyanophycese, and in Gloeo- 

 trichia and many other Rivulariacese they are a constant fea- 

 ture. (5) The granular bodies which appear in the Cyano- 

 phycean protoplast, are always observed outside the 'cen- 

 tral body.' According to their reactions, they are divided 

 into two widely different groups, the 'cyanophycin gran- 

 ules' and the 'slime balls.' The cyanophycin granules, 

 which under ordinary conditions apparently consist of a 

 firm substance, are easily soluble in dilute hydrochloric acid, 

 are colored blue by haematoxylin, and when stained intra- 

 vitam, retain no methyl blue. They are generally found in 

 the outermost part of the periphery of the chromatophore, 

 and are doubtless to be regarded as the first visible assimila- 

 tion product of the activity of the chromatophore. In the 

 spores are placed the necessary reserve materials for their 

 germination. The slime balls, which are composed of a 

 tenacious, flowing substance, are insoluble in dilute hydro- 

 chloric acid, are colored red-violet with haematoxylin, and 

 store up methyl blue during intra-vitam staining. They 

 are closely placed around the central body, and only occa- 

 sionally appear further out in the chromatophore. Their 

 significance in the cell is not clear. The 'nucleolus,' the 

 'central substance' and the 'red granules' of other authors 

 are bodies identical with the slime balls. (6) In the germ- 

 ination of Gloeotrichia spores, oil appears in the chromato- 

 phores of the cells. Zukal had also noticed this in other 

 Cyanophycese. It can be established conclusively that the 

 Cyanophycean protoplast is made up of at least two different 

 parts, sharply distinguished from one another, and it is ques- 

 tionable whether we should call the central body a cell 

 nucleus." Following this he gave a somewhat lengthy dis- 

 cussion of the phylogenetic relationship of the central body 

 to the cell nucleus, and he concluded that on account of the 

 "lack of a chromatin framework, the absence of nuclei, the 

 direct division which may be more complicated than it 



