Structure, etc., of Epiphegus Virginiana. 377 



bundles are relatively well developed. There is a core of long 

 slender spiral tracheae extending out farthest in the lobes. 

 On both sides of this occur short stout spiral tracheids, set 

 irregularly and discontinuously along the bundle. A few 

 small phloem cells lie to one side. The tissue cells are much 

 larger than the bundle cells. They fit together in all kinds 

 of positions, except near the bundles, where they arrange 

 themselves quite regularly. 



One-celled, rarely two-celled hairs fringe the edges of the 

 lobes. Below, across the base of the lobes, there extends 

 a band of two or three-celled hairs, longer than the upper 

 hairs. All of these hairs are on the outer surface of the 

 calyx ; none are present on the inner surface. They have a 

 swollen granular appearance. 



The Corolla. — In the chasmogamic flower (Fig. 3) the 

 corolla consists of cells longitudinally elongated, and is of 

 four or three cells in thickness. The outer layer of cells 

 is quite regular and even, constituting an epidermis. At 

 the base, where the corolla passes under the calyx, it 

 thickens greatly, being many times thicker than either the 

 calyx or the upper corolla region, and consisting of two cell 

 layers. Five vascular bundles pass through its length and 

 end in the five corolla lobes. They are similar to the bundles 

 of the calyx. 



There are two areas of hair development, both on the 

 inner surface. One area forms a hairy belt along the outer 

 margins of the corolla lobes. These are short, mostly two- 

 celled hairs, and often so bend on themselves as to have a 

 hooked appearance. They are greatly swollen and turgid- 

 looking. The other area is a ring extending round the inner 

 surface at about the region where the stamens are inserted. 

 These hairs are much longer than those above. They con- 

 sist of about three narrow elongated cells. At the free end 

 of each hair there is often a flat, flange-like expansion. 



The cleistogamic corolla is greatly reduced in size. It 

 becomes an apparently closed asymmetric cap-like structure 



