378 Cooke and Schively on Observations on the 



that is lifted off by the maturing ovary. The bundles are 

 also reduced. A small fifth one may appear, but usually 

 there are but four bundles. Two of these are fairly large, 

 and show both spiral tracheae in three or four rows, and 

 tracheids. They occur about one-third of the circumference 

 of the corolla apart, and arise on the anterior short curve 

 of the cap-like corolla. In that position they can best sup- 

 port the corolla. The two small bundles alternate with these 

 large ones. They apparently contain but a single spiral 

 trachea, that extends clear to the end of the corolla. The 

 larger bundles end some distance below. Stomata have 

 been found on the corolla. 



This corolla is from four to six cells in thickness. The 

 outer layer is of large regular close-set cells, with a con- 

 siderable amount of thickening developed on their outer 

 walls. A protective epidermis is thus supplied to it. Longi- 

 tudinal sections reveal that the corolla is not in strict mor- 

 phological sense a cleistogamic one, but has a distinct 

 though small mouth (Fig. 4). This opening occurs where 

 the anterior straight, or slightly concave and convex sur- 

 faces of the corolla meet. The convex surface develops a 

 disc of greatly thickened cells that overlaps the concave 

 side, leaving a distinct mouth. This mouth is entirely closed 

 by the numerous small hairs crowding into it from within 

 the corolla. These hairs are of one to two round swollen 

 cells. They arise in great numbers on the inner surface 

 of the corolla all about this mouth. Also a group of hairs 

 appears on the outer surface of the corolla, tip, just where it 

 is overlapped by the other surface. These hairs help greatly 

 in filling up this mouth. The corolla may then be called 

 physiologically but not morphologically cleistogamic. There 

 is a slight development of long hairs about the region where 

 the stamens arise. 



The hairs closing the corolla's mouth are evidently in- 

 tended to keep out intruding insects. Probably the numer- 

 ous hairs developed on the chasmogamic flowers have also a 



