158 CARROLL— ON DEVELOPMENT OF 



formed of two joined, and the corolla as composed of four petals joined 

 in pairs, the fifth being aborted. Bernhardi viewed the two small 

 green sepals as two bracts. The calyx to him was composed of five 

 members — an inferior spurred sepal, a superior sepal composed of two 

 joined, and two little sepals which usually aborted and which botanists 

 had not mentioned. The corolla was composed of five petals — four 

 joined in pairs and a fifth which had united with the two joined superior 

 sepals. Roeper viewed the calyx as composed of five parts — two were 

 often rudimentary or disappeared, two were small lateral sepals, 

 and the fifth spurred sepal was toward the axis of the inflorescence. The 

 corolla was composed of five petals which alternated with the sepals — 

 four were joined in pairs, the fifth was the superior petal of the older 

 botanists. Payer accepted the view of Roeper, and agreed that the two 

 small prominences representing the anterior sepals were to be found 

 in the young stage of Impatiens balsamina and figured them in the 

 young stages of Impatiens Royleana. Lindley (28) followed the view of 

 Knuth. Henfrey (22) arguing from the morphology of double balsams 

 adopted Roeper's view although he failed to find the rudiments of the 

 two anterior sepals reported by Roeper and Payer in Impatiens bal- 

 samina. The view of Roeper was adopted by Warburg and Reiche 

 in " Pflanzenfamilien. " Gray's (18) view agreed with that of Knuth, 

 and this was reported in the seventh edition of Gray's Manual. Brit- 

 ton and Brown's Manual followed the view of Roeper. The double 

 appearance of the anterior member in our native species would seem 

 to suggest that it is formed of two members which if it be regarded 

 as belonging to the petaline whorl would raise the number to six mem- 

 bers, but if it be regarded as belonging to the sepaline whorl would 

 raise the number to five. The occurrence occasionally of two spurs 

 upon this structure would further emphasize its double nature. The 

 fact that supernumerary spurs occur upon the two lateral usually 

 greenish sepals, but not upon the two-lobed petals would tend to argue 

 its connection with the sepaline whorl. The occurrence in other species 

 of the genus of spurs on all five structures further argues for this 

 view. In this paper, I have therefore adopted provisionally the view 

 of Knuth and Gray in describing the calyx as composed of four sepals, 

 the anterior sepal being formed of two fused sepals and the corolla 

 as composed of two-lobed petals representing four petals fused in pairs. 

 These have been well figured by Gray (18). 



Peloria is not uncommon in the genus Impatiens. It is stated that 

 the two lateral sepals are often spurred in double garden balsams. 



