412 RUSSELL— A COMPARATIVE STUDY OF 



tion between the carpellary walls and a broad zone of large thin-walled 

 cells. This is shown, by its reaction to iodine, to be a starch storing 

 layer. In its outer part, lying nearer to the carpellary wall, are numerous 

 small vascular bundles, cut transversely. This zone is, from all evidence 

 at hand, a wide development of the tissue of the integuments surrounding 

 the single ovule. The wide development and apparent fusion of the in- 

 teguments, and the apparent single layer of the nucellar tissue, breaking 

 down early in the development of the ovule, are characteristics resembling 

 the development of the ovules of Impatiens fulva, as described by Mr. 

 Franklin Carroll in the present publication ("The Development of the 

 Chasmogamous and the Cleistogamous Flowers of Impatiens fulva"). 

 Floerkea represents perhaps a more reduced state than that found in 

 Impatiens fulva. Further careful study of the development of the ovule 

 of Floerkea would be necessary, however, before a definite opinion on the 

 matter could be expressed. 



Within the fused integuments in the more nearly matured seed is a 

 broken down granular substance, immediately surrounding the remnants 

 of the embryo sac membrane. This represents the broken down nuceUar 

 tissue. Within the embryo sac is the embryo showmg the cotyledons 

 cut transversely, and the radicle between. In the spaces about the 

 maturing embryo are shreds of the broken down endosperm tissue almost 

 entirely consumed by the embryo at this stage of growth. 



In L. Douglasii the flowers are solitary axillary. The flowers are 

 5-merous in symmetry, having 5 sepals, valvate in aestivation, green and 

 persistent, united at the base as in the F. proserpinacoides. The petals 

 are spatulate and about twice the length of the sepals. They are whitish 

 or yellowish in color. On the base of the petal are unicellular hairs of 

 the same type as in Floerkea, but longer. Fig. 10 gives the relative size 

 of the hairs in Floerkea and Limnanthes. On the middle regions of 

 the petals in Limnanthes, however, are hairs not present on the 

 petals of F. proserpinacoides. These are longer than the petals of 

 F. proserpinacoides and are shown in relative size in Fig. 10. These 

 hairs are peculiar in their shape and walls. They are unicellular, and 

 tapering. Their walls are covered with wart-hke thickenings resembhng 

 the thickenings of the walls of the epidermis in the carpels of Floerkea 

 and Limnanthes. 



The stamens are 10, in two rows. The outer row, opposite the sepals, 

 and not the row opposite to the petals, as is stated in some descriptions, 

 are the ones which bear the gland, as in Floerkea. The stamens are 



