329 BOTANICAL GAZETTE [april 



appearance. They originate from the immediate segments of the 

 apical cell, and their development is strictly acropetal. 



4. The antheridia develop according to the usual method 

 found among the anacrogynous Jungermanniales. Variations 

 occur in the order of appearance of the walls in the primary stalk 

 cell. 



5. Until the appearance of the first vertical wall, young arche- 

 gonia cannot be distinguished from young antheridia. The first 

 transverse division in the archegonium initial separates the stalk 

 cell from the archegonium proper, and subsequent development 

 follows the usual Jungermanniales type. The cover cell is inactive, 

 6-8 neck canal cells are formed, and the venter is 2 cells thick be- 

 fore fertilization. The archegonium is of an advanced type. 



6. The early divisions of the embryo are transverse, both 

 halves of the fertilized egg contributing to the development of 

 the foot, seta, and capsule. A calyptra 3-4 cells in thickness is 

 formed. 



7. The sporogenous tissue is differentiated rather early in the 

 history of the sporophyte. The elaters are rudimentary, and 

 each is homologous with a single spore mother cell, not with a row 

 of them. 



8. The sporophyte is primitive. 



To Dr. W. J. G. Land, under whose direction the study was 

 made, the writer makes grateful acknowledgment for his kind 

 advice and helpful criticism. 



Carthage College 

 Carthage, III. 



LITERATURE CITED 



1. Austin, Coe F., Characters of some new Hepaticae. Proc. Acad. Nat. Sci. 

 Philadelphia, p. 228. 1869. 



2. Cavers, F., The interrelationships of the Bryophyta. III. Anacrogynous 

 Jungermanniales. New Phytol. 9:197-234. 1910. 



3. Evans, A. W., Notes on New England Hepaticae. XII. Rhodora 17:107- 

 iii. 1915. 



4. Haupt, A. W., A morphological study of Pallavicinia Lyellii. Bot.Gaz. 

 66:524-533. 1918. 



5. Hill, E. J., Fossombronia crispula in the dune region of Indiana. Bryolo- 

 gist 19:67-68. 1916. 



