78 Fertility in Cicliorium intybus 



of the first self-fertile parent (^1 x ^22) no. 4 wfis very low, being only 

 4 7o- Although 7 of the 10 plants grown from its seed Avere self-fertile, 

 the highest individual fertility was 237o> and the average was 14 7o- 

 The four plants selected from this series as parents had fertilities of 12, 

 13, 16, and 22. The numbers of plants grown from these were too 

 small for an adequate judgment of the various series, but from the two 

 most highly self-fertile plants were derived the series (A y.E'22)-4-3-ll- 

 and {A x E,i2)-.'f.-6-3- reported above. The first self-fertile parent was 

 very feebly self-fertile, but with one exception parents for subsequent 

 series were selected for highest individual fertility. 



The behaviour of the various series indicates an irregular inheritance 

 of the characteristics of self-compatibility and the frequent and decided 

 regression to self-sterility. The range of the degi'ee of self-fertility was 

 rather decidedly extended in the series {A x E2'2)-4-6-3-, and the 

 average fertility was also high. This series was decidedly more fertile 

 in every comparison than the series {A x E22)-4-3-ll~ : the ancestral 

 fertilities were only slightly higher, being 4, 22, and 31 as compared 

 with 4, 13, and 32. 



Considered as a whole, the family {A x E22)-4- had 40 "/^ of the 

 total plants self-fertile. The distribution on the basis of fertilities of 

 the self-fertile plants is irregular and skew, the percentage of self- 

 fertility observed in an individual is extended to G2, and the average 

 percentage for all self- fertile plants is 16'5. 



{h) The family {A x E22)-9-. 



As indicated in Table VII, the third filial generation in this family 

 consisted of six series, which were derived from as many different plants 

 of the second generation which had in turn descended from two plants 

 of the first generation. Thus the series are of the two main sub- 

 families {A X E22)-9~-4- and {A x E22)-9-5-. 



The data for the series which descended from plant {A x E22)-9- no. 4- 

 are given in Table III. Of the 10 plants in series {A x E22)-9-4-4.- , 

 four were self- fertile ; of 13 in series {A x E22)-9-4-10- , seven were 

 self-fertile; and of the 4 plants in series {A x E22)-9-4-ll-, two were 

 self-fertile. The fertilities of the immediate parentage were quite alike, 

 the complete record of ancestry being 23, 43, and 20 for the first 

 mentioned, 23, 43, and 1 7 for the second, and 23, 43 and 20 for the last 

 named. As shown in the tables, the proportional number of self-fertile 

 plants varies, as do the individual fertilities. The number of plants is 

 perhaps insufficient for adequate comparisons of differences. It is to be 



