— 23 — 



perianths and bracts. Ingham,* however, lists it without question from West 

 l^ncashire and from South Aberdeen. Specimens from the original locality 

 and from Koppelsberg in Pomerania have been distributed by Schiffner in his 

 Hep. Europ. Exsic, Nos. 543 and 544. 



The group of Cephaloziae in which the leaves have more or less connivent 

 lobes includes a number of closely related species. Of this group C. catenulata 

 (Hiihen) Dum., C. connivens (Dicks.) Lindb., C. media Lindb., and C. pleniceps 

 (Aust.) Lindb. have long been known in both Europe and North America, while 

 C. affinis Lindb., C. compacta VVarnst., C. Loitlesbergeri SchifFn., and C. macro- 

 stachya Kaalaas have been published during the past few years. These recent 

 segregates were based on European material, but C. affinis"- and C. macrostachya' 

 have since been definitely reported from North America. Now that C. Loitles- 

 hergeri has been added to our flora, the discovery of C. compacta would not be 

 surprising. 



The essential characters of C. Loitlesbergeri may be briefly described as 

 follows: the color is pale or yellowish green; the lobes of the leaves are long, 

 entire, and usually connivent and acuminate, each being tipped by a row of from 

 two to four cells; the leaf-cells have firm walls without trigones and average about 

 30/U in diameter; the inflorescence is autoicous; the innermost perichaetial bracts 

 are deeply divided into four or more, sub-parallel and long-acuminate lobes 

 separated by narrow sinuses; the mouth of the perianth is long-ciliate. Although 

 the lobes of the bracts are sometimes subequal in size, the two median lobes are 

 often longer and broader than the others, so that the lobes might be described 

 as bifid with accessory lobes. 



It will at once be seen that the essential characters of C. Loitlesbergeri are 

 shared by other members of the group to which it belongs, although they occur 

 in diff'erent combinations. In C connivens, for example, the inflorescence is 

 autoicous, the lobes of the bracts are numerous and long-pointed, and the perianth 

 is ciliate at the mouth; in C. affinis the inflorescence is autoicous, and the mouth 

 of the perianth is ciliate; in both C. macrostachya and C. catenulata the perianth 

 is long-ciliate; and in C. pleniceps the inflorescence is autoicous. In C. connivens, 

 however, the cells are much larger than in C. Loitlesbergeri, averaging 50^ or 

 more in diameter; in C. affinis the lobes of the bracts are broader and blunter, 

 and the cilia at the mouth of the perianth are shorter; in C. macrostachya the 

 inflorescence is dioicous and the bracts are distinctly bifid with shorter lobes; 

 in C. catenulata the inflorescence is dioicous, and the shortly bifid bracts are 

 coarsely dentate; in C. pleniceps the lobes of the bracts, although sometimes more 

 than two, are broader and acute, rather than acuminate, while the mouth of 

 the perianth is crenulate. In C. media, v^Wich. often grows with C. Loitlesbergeri, 

 the inflorescence is dioicous, the bracts are distinctly bifid (although sometimes 

 with supplementary teeth), and the mouth of the perianth is crenulate. 



I 



1 Census Cat. Brit. Hepat. 26. 1913- 



= See Evans, Bryologist 17: 89. 1914- 



' See Schiffner. Hedwigia 54 : 322. 1914; also Evans, Rhodora 17 : 1 14- I9IS. 



