MORPHOLOGICAL REVISION 139 



of these is the smallest, articulating between the tibiale and the first distal, 

 and, on the outer side, with another centrale. This is an unusual position 

 for a centrale, and I have endeavored to find in the small bone some evi- 

 dence of extraneous origin, but am quite convinced that it really belongs 

 in this place, as otherwise the space it occupies must have been unossified. 

 The median distal centrale is the largest of the three distal centralia, and 

 is nearly square in outline; it articulates proximally with the tibiale and 

 proximal centrale, on either side with a centrale, and distally with the second 

 distal. Of the distalia, the second is the largest and the fifth is the smallest, 

 the third and fourth smaller than the second. Each distal supports its own 

 digit exclusively. 



"The digits are, it is seen by the figure and the restoration, very short 

 and heavy. The first metatarsal is very characteristic in its broad and short 

 form, resembling more a proximal phalanx, broadly expanded proximally 

 and much constricted in the middle. At its distal extremity there is a 

 fragment of the first phalanx in articulation, the remainder being lost. 

 The width of this fragment would indicate the possibility of a second pha- 

 lanx of very small size. The second metatarsal is much longer than the 

 first, its proximal extremity less expanded. Its first phalanx is short and 

 expanded, much shorter and smaller than the corresponding phalanx of 

 the third finger. At its distal extremity is preserved the proximal extremity 

 of a very small second phalanx, and there was probably no more present 

 in the living animal. The third metatarsal is much longer than the second, 

 with its proximal extremity more oblique to its long axis. Its distal extremity 

 is quite on a line with the distal extremity of the second metatarsal. The 

 first phalanx is stouter and longer than the first phalanx of the second 

 digit, and a trifle smaller than that of the fourth, its greatest width about 

 equal to two-thirds its length. The second phalanx, preserved entire, is 

 much smaller than the first, and is very short and broad, its width equal 

 to its length; it is but little constricted at its middle. A proximal half or 

 end of a minute third phalanx is also present. It was somewhat pointed 

 in shape, but by no means a claw. The fourth metatarsal is much like the 

 third and of about the same length, its proximal extremity yet more oblique. 

 Its proximal phalanx is a little longer than the first of the third digit, its 

 distal extremity surpassing a little the distal extremity of the third meta- 

 tarsal. The second phalanx is a little longer but no broader than the sec- 

 ond of the third toe. Its third phalanx is about two-thirds the length of 

 the second, but is much narrower distally. The basal part of the fourth 

 phalanx, a very small bone, is articulated with the third, and it is very evi- 

 dent that it was the terminal one, and was very small and in no sense a 

 claw. The fifth toe is slender, and was divaricated in life. In the specimen, 

 while still retaining its articulation with the tarsus, it is turned across the 

 fourth and third metatarsals. Its metatarsal is much narrower, and not 

 more than three-fourths the length of the fourth. Its first phalanx, likewise 

 slender, is about the length of the first phalanx of the second toe, but is 

 much narrower. The second phalanx, much shorter, is narrowed distally, 

 but with a minute terminal knob or expansion. Possibly an ossicle not 

 larger than a pin-head may have articulated here, but probably not. 



"The actual phalangeal formula of the foot as preserved is, it is seen, 

 I, 2, 3, 4, 2. It is possible, though not very probable, that the first, second, 

 and fifth digits may have had a minute ossicle at the extremity of each, 



