294 BOTANICAL GAZETTE [October 



the growth on the dorsal side of the sporophyll is very marked, 

 resulting in the deflexing of the coalesced bract and scale. The 

 bract is free only at the tip of the deflexed portion. The vascular 

 systems to the bract and scale are distinct from the strobilus cylin- 

 der. The bract bundle remains single to the tip of the bract. The 

 two scale bundles end in a dorsal row of numerous inverted bundles. 

 The flank bundle may turn partly and lie nearer the ventral side 

 of the appendage. 



The strobilus of Juniperus communis (figs. 105-113) consists of 

 a whorl of three sporophylls united at the base and surrounding 

 three ovules. In the older stage the sporophylls coalesce also at 

 the tip, forming a fleshy berry-like body. The coalescence of 

 bract and scale is complete. 



After the traces to the three bracts beneath the sporophylls 

 have left the cylinder, each of the three remaining bundles divides 

 into three; the median or bract bundle proceeds undivided into 

 the bract; the two lateral or scale bundles divide, giving off some 

 bundles to the dorsal side of sporophyll and some to ventral in 

 such way that, including bract bundle, an oval ring of bundles 

 with xylem facing is formed. 



Taxodineae 



The general features of the ovulate strobilus in the Taxodineae 

 are spiral arrangement of sporophylls, reduction in number of 

 sporophylls in some forms, considerable coalescence of bract and 

 scale, and varying number and orientation of ovules. 



In Cryptomeria japonica (figs. 1 14-128) the axis of the young 

 strobilus is very short and the sporophylls are crowded at the 

 broadened summit. The ovules at this stage appear to be inserted 

 on the strobilus axis. The scale begins its development as four to 

 six lobes between the ovules and the dorsal side of the bract. In 

 an older stage the axis has elongated, forming a globose cone. The 

 erect ovules, usually three in number, are definitely inserted on 

 the sporophyll. The lobed scale is united for two-thirds of its 

 length to the bract. In the upper portion of the cone three bundles 

 leave the axis for the sporophyll; one at the base of the gap to 

 supply the bract, and one from each side of gap to supply the scale. 



