370 BOTANICAL GAZETTE [November 



fall after blooming, but in some varieties they persist even after 

 the ripening of the fruit. 



The staminate flower begins to develop as early as the pistillate, 

 but the pistils develop more rapidly than the stamens, resulting 

 in the sterilization of stamens and in some cases in their disap- 

 pearance. 



The mature pistillate flower generally has two sets of stamens 

 (nos. i and 2), but in some cases has more (nos. 1,2, and some of 

 4) or less (no. 1 and some of no. 2). 



I did not discover the following condition described by Miss 

 Hague: "In the pistillate flower it is a common occurrence to 

 rind the number increased by the branching of one or more of the 

 stamens." 



In the development of the pistil there appear first four large 

 protrusions, and then four smaller ones between them (figs. 5 and 

 6, a, b). The larger ones give rise to two small papillae, which are 

 the initials of ovules (figs. 6, d, and 7). The papillae grow at first 

 toward the wall of the ovary, then curve downward toward the 

 center, and finally upward (figs. 8 and 9). Simultaneously the 

 small protrusions develop toward the center of the ovary, resulting, 

 along with the development of larger protrusions, in 8 loculi for the 

 8 ovules. 



The ovule has two integuments (fig. 9), which extend beyond the 

 nucellus like a beak at the time the embryo sac is fully developed 

 (fig. 10). 



Megaspore formation 



In all the varieties examined the archesporial cell is solitary 

 and hypodermal. There is no division resulting in a parietal cell, 

 so that the archesporial cell is the megaspore mother cell. The 

 heterotypic division of this nucleus occurs in the same way as in 

 the case of the nucleus of the microspore mother cell. After 

 nuclear division, the cytoplasm organizes into two parts, the inner 

 one being the larger. Sometimes a wall between these two parts 

 does not appear before another division of nuclei occurs. 



Wall formation in connection with the megaspores is variable 

 in direction. Sometimes the two walls are perpendicular to one 



