11 



The crinkled-stamen type of flower was found in each one of 34 

 hui the length of filaments and the <\<^v<'M of crinkling 

 differed somewhat as La shown by comparing Figs. 22 and 23. In 

 1920 all but 2 of the 34 plants produced seeded fruit and of tb 

 bore a " full " crop, 9 a " medium " crop, 9a" light " crop, and 7 

 a "very light" crop. With the exception of two plant-, these 

 seedlings were able to function as females in fruit and seed pro- 

 duction. 



The ancestry of this series of 34 plants is well known for three 

 generations. First, Winchell was crossed with Diamond (both 

 classed as hermaphrodites with upright stamens), and Station 

 Seedling No. 123 with upright stamens was obtained. This seedling 

 was used as a pollen parent on Worden, also with upright stamens, 

 and three seedlings, all with upright stamens, Xos. 931, 033, and 934, 

 were obtained. The 34 plants derived from selling seedling No. 933 

 were all of the type with crinkled stamens. Of the progeny obtained 

 by selfing the sister plants, 931 and 934, 24 have bloomed and 

 have only the normal erect stamens of the perfect hermaphrodite. 

 As to length, the stamens of 1 were; short, of 20 medium, and of 3 

 long. 



It appears that the plants with this crinkled type of stamen are 

 not able to function as males. Functionally their sex is quite like 

 that of the imperfect hermaphrodites with reflexed stamens. One 

 might consider that in these there is a combination of the " reflexed " 

 character of filament with the " upright." The appearance of this 

 type of flower in all of 34 sister plants after three generations of 

 parentage with normal upright stamens suggests the phenomenon 

 of the so-called mass mutation. 



Another plant was found bearing flowers with crinkled but less 

 erect stamens (Fig. 26). In this case the filaments may be con- 

 sidered as tending to be recurved. This plant is a seedling derived 

 from selfing Station Seedling No. 4574 (having normal upright 

 stamens), which was an offspring of Salem (reflexed) crossed with 

 Worsen (upright). In this case the reflexed character was pre 

 in one grandparent. 



Whether the crinkled character of stamen is to be considered as 

 due to a recombination of hereditary units, as an intermediate 



