V YORK 

 A NIC At. 

 UAKDfiM 



Reprinted from the American Journal of Botany, X: 485-498, November, 1923. 



GENERA OF NORTH AMERICAN FABACEAE 

 I. TRIBE GALEGEAE 



P. A. Rydberg 

 (Received for publication December 16, 1922) 



The tribe Galegeae has been divided since Bentham's time into seven 

 subtribes. Of these one has already been raised to tribal rank, Psoralieae, 

 which is characterized by the glandular-punctate foliage and by the pods, 

 which are one- or few-seeded, usually indehiscent but rarely breaking open 

 irregularly across the middle, never valvate. Another subtribe should be 

 removed as a tribe Indigofereae, having three characters seldom found 

 elsewhere in Fabaceae and never combined in any of the tribes. These are 

 malpighiaceous hairs (appressed hairs, 2-branched with branches forming a 

 straight line, the hairs therefore apparently attached at their middle), 

 appendaged connectives in the anthers, and lateral spurs on the keel-petals. 

 It may be that Glycyrrhiza and one or two genera should be removed also 

 and form another tribe, on account of the indehiscent (though several- 

 seeded) glandular fruit and confluent anther cells. 



After the removal of the two subtribes mentioned, the tribe may be 

 characterized as follows: 



Herbs, shrubs, or trees, or woody vines, without glandular dotted 

 foliage (except in Glycyrrhiza). The leaves are odd-pinnate or abruptly 

 pinnate, with or without stipels, very rarely with malpighiaceous hairs. 

 The calyx is campanulate to tubular, 5-toothed or 5-lobed, rarely more or 

 less 2-tipped. The corolla is papilionaceous, with 5 petals, the two keel- 

 petals more or less adnate along the lower margins. The stamens are 10, 

 diadelphous or monadelphous, the connective without appendages. The 

 ovary is usually many-ovuled, more or less curved or bent upwards; the 

 style slender and the stigma small. The fruit is several- or many-seeded, 

 2-valved, usually dehiscent. 



As stated before, Bentham and Hooker acknowledged 7 subtribes. 

 These authors were closely followed by Taubert in Engler and Prantl's 

 Natiirlichen Pfanzenfamilien. Taubert's key in translation reads as 

 follows, after the Indigofereae and Psoralieae have been removed: 



a. Seeds strophiolate. Flowers in 2's in the leaf-axils or forming a 



terminal raceme (compare Tephrosia and Fordia). c. Brongniartiinae. 



/3. Seeds not strophiolate. 



I. Inflorescence racemose, terminal, opposite the leaves, or on 

 the branches paniculate, more rarely in the axils of the 

 upper leaves or all peduncles or only the lower paired or 

 clustered in the axils. Stipe inside the staminal sheath 

 sometimes surrounded by a small cup-like disk. d. Tephrosiinae. 



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