﻿APR. 
  19, 
  1922 
  HITCHCOCK 
  : 
  PERENNIAL 
  SPECIES 
  OF 
  TEOSINTE; 
  207 
  

  

  near 
  Olyra 
  but 
  he 
  thinks 
  it 
  stands 
  near 
  Zea 
  (Indian 
  corn), 
  in 
  fact, 
  that 
  it 
  

   resembles 
  closely 
  a 
  stunted 
  plant 
  of 
  maize. 
  He 
  points 
  out 
  that 
  the 
  staminate 
  

   spikelets 
  are 
  2-flowered 
  instead 
  of 
  1 
  -flowered 
  as 
  described 
  by 
  Schrader; 
  

   describes 
  fullv 
  the 
  female 
  or 
  pistillate 
  spikelets 
  and 
  discusses 
  the 
  relation 
  to 
  

   Tripsacum 
  and 
  Zea, 
  stating 
  that 
  Euchlaena 
  is 
  a 
  Zea 
  in 
  which 
  the 
  female 
  

   inflorescence 
  is 
  nearly 
  as 
  in 
  Tripsacum; 
  and 
  quotes 
  Grisbach 
  (Veg. 
  Erde 
  1: 
  

   542) 
  as 
  doubting 
  the 
  American 
  origin 
  of 
  corn 
  because 
  of 
  its 
  affinity 
  with 
  certain 
  

   Asiatic 
  genera 
  such 
  as 
  Coix, 
  but 
  Ascherson 
  himself 
  thinks 
  Zea 
  is 
  much 
  more 
  

   closely 
  related 
  to 
  Tripsacum, 
  an 
  American 
  genus. 
  Ascherson 
  discusses 
  

   Reana 
  luxurians 
  and 
  takes 
  occasion 
  to 
  transfer 
  it 
  to 
  Euchlaena, 
  of 
  which 
  genus 
  

   he 
  considers 
  it 
  a 
  second 
  species 
  differing 
  in 
  its 
  greater 
  size. 
  There 
  are 
  as 
  

   manv 
  as 
  150 
  culms 
  to 
  one 
  plant, 
  these 
  being 
  as 
  much 
  as 
  2V2 
  meters 
  tall. 
  

   In 
  the 
  only 
  staminate 
  spikelet 
  of 
  E. 
  mexicana 
  he 
  has 
  seen 
  the 
  lemmas 
  are 
  

   shorter 
  than 
  the 
  glumes 
  while 
  in 
  E. 
  luxurians 
  they 
  are 
  as 
  long 
  as 
  the 
  

   glumes. 
  The 
  joints 
  of 
  the 
  pistillate 
  inflorescence 
  are 
  cylindrical 
  and 
  ob- 
  

   liquely 
  truncated 
  at 
  the 
  ends 
  instead 
  of 
  being 
  triangular 
  as 
  in 
  E. 
  mexicana. 
  

   This 
  is 
  the 
  same 
  difference 
  distinguishing 
  the 
  Florida 
  form 
  of 
  the 
  cultivated 
  

   teosinte 
  from 
  the 
  Dmrango 
  form 
  as 
  pointed 
  out 
  by 
  Mr. 
  Collins. 
  

  

  In 
  1910 
  I 
  collected 
  in 
  Mexico, 
  near 
  Zapotlan, 
  now 
  called 
  Ciudad 
  

   Guzman, 
  a 
  perennial 
  species 
  of 
  Euchlaena, 
  and 
  Mr. 
  G. 
  N. 
  Collins 
  

   collected 
  it 
  at 
  the 
  same 
  place 
  in 
  October, 
  1921, 
  while 
  searching 
  for 
  

   teosintes 
  in 
  their 
  native 
  habitat. 
  This 
  species 
  differs 
  distinctly 
  from 
  

   all 
  previously 
  known 
  forms 
  of 
  teosinte 
  in 
  the 
  possession 
  of 
  rhizomes 
  and 
  

   is 
  described 
  below 
  as 
  new. 
  

  

  Euchlaena 
  perennis 
  Hitchc, 
  sp. 
  nov. 
  

  

  Plants 
  perennial, 
  producing 
  strong 
  scaly 
  rhizomes; 
  culms 
  erect 
  or 
  somewhat 
  

   geniculate 
  at 
  base, 
  firm, 
  glabrous, 
  1 
  to 
  2 
  meters 
  tall; 
  sheaths 
  striate, 
  the 
  striae 
  

   joined 
  by 
  numerous 
  cross-veins, 
  glabrous 
  or 
  some 
  of 
  them, 
  especially 
  the 
  upper 
  

   or 
  those 
  of 
  the 
  branches, 
  somewhat 
  hispid 
  in 
  the 
  region 
  of 
  the 
  collar 
  and 
  

   throat, 
  the 
  lower 
  longer 
  than 
  the 
  internodes, 
  the 
  upper 
  shorter; 
  ligule 
  a 
  short 
  

   somewhat 
  lacerate 
  membrane, 
  1 
  to 
  2 
  mm. 
  long; 
  blades 
  linear 
  or 
  linear- 
  

   lanceolate, 
  as 
  much 
  as 
  40 
  cm. 
  long 
  and 
  3 
  cm. 
  wide, 
  the 
  upper 
  shorter, 
  some- 
  

   what 
  cordate-clasping 
  at 
  base, 
  acuminate, 
  flat 
  and 
  rather 
  thin, 
  the 
  white 
  

   midnerve 
  prominent 
  beneath, 
  glabrous, 
  strongly 
  scabrous 
  or 
  scabrous-ciliate 
  

   on 
  the 
  margin, 
  ciliate 
  near 
  the 
  base; 
  terminal 
  inflorescence 
  staminate, 
  con- 
  

   sisting 
  of 
  2 
  to 
  5 
  approximate, 
  ascending 
  or 
  spreading 
  racemes 
  6 
  to 
  12 
  cm. 
  long, 
  

   the 
  internodes 
  between 
  the 
  lower 
  ones 
  about 
  1 
  cm. 
  ; 
  spikelets 
  in 
  pairs, 
  the 
  pairs 
  

   alternately 
  to 
  right 
  and 
  left 
  on 
  one 
  side 
  of 
  a 
  flat-triangular 
  rachis, 
  the 
  rachis 
  

   internodes 
  5 
  to 
  8 
  mm. 
  long, 
  scaberulous 
  or 
  ciliate 
  on 
  the 
  angles; 
  spikelets 
  2- 
  

   flowered, 
  8 
  to 
  9 
  mm. 
  long, 
  elliptic 
  or 
  somewhat 
  broader 
  above, 
  the 
  middle 
  one 
  

   of 
  the 
  pair 
  nearly 
  sessile, 
  the 
  other 
  on 
  an 
  angular 
  scaberulous 
  pedicel 
  3 
  to 
  4 
  mm. 
  

   long, 
  enlarged 
  toward 
  apex; 
  first 
  glume 
  flat 
  on 
  the 
  back, 
  strongly 
  inflexed 
  at 
  

   the 
  margins, 
  smooth 
  except 
  the 
  scaberulous-ciliate 
  keels, 
  these 
  somewhat 
  

   winged 
  above, 
  slightly 
  notched 
  at 
  apex, 
  the 
  midnerve 
  rather 
  faint, 
  the 
  strong 
  

   lateral 
  nerves 
  at 
  the 
  inflexed 
  margins, 
  a 
  second 
  faint 
  pair 
  intermediate; 
  

  

  