38 ILLINOIS BIOLOGICAL MONOGRAPHS [388 



130/*. The ovary is distinctly, but not considerably, to the pore side of 

 the median line. It is fanshaped in outline, being formed of a large 

 central mass from which about 30 lobes radiate extending in all direc- 

 tions in the horizontal plane except towards the part of the field occu- 

 pied by the vitelline gland. It is distinctly ventral in position. In 

 two specimens however most of the proglottids which have the genital 

 pore on the left side have the ovary separated into distinct dorsal and 

 ventral portions with a narrow connection between them. The dorsal 

 portion in these cases is of about half the area of the ventral and thin. 

 These are the individuals that show the peculiarities of the excretory 

 ducts already mentioned. The oviduct in all cases connects with the 

 ovary at a point directly in front of the vitelline gland and is short. 

 The vitelline gland is of the ordinary bilobed type which is found nearly 

 everywhere in the subfamily and shows no suggestion of the type found 

 usually in the genus. It lies posteriorly to the ovary, sometimes ex- 

 tending dorsad to the cortical layer and dividing the testes into two 

 fields. In cross section it is distinctly horseshoeshaped, with the open 

 face dorsad. Viewed from above it is U-shaped but with the median 

 arm much larger. 



The uterus is first recognizable as a sheet of deeply staining cells 

 ventrad of the testes and dorsad of the ovary. Just beneath the mar- 

 gin of each testicular field the sheet is thickened into a heavy circular 

 band. On the pore side there extends forward from the anterior tip 

 of the circular band a strip of the same nature, which crosses the vagina 

 laterad of the receptaculum seminis then turns and extends diagonally 

 distad just underneath the anterior margin of the receptaculum and 

 anterior to the vitelline gland. It joins with the other circular band at 

 the corner of this gland. It is into this cross duct that the uterine duct 

 empties. Within these circles and anterior to the transverse portion 

 and connecting with them at frequent points, is a network of strands 

 formed by thickenings of the uterine tissue which recalls strongly the 

 reticulate uterus found in Moniezia. The transverse portion is of course 

 the first part of the uterus to receive eggs; but the eggs pass immedi- 

 ately into the circular bands which have developed into canals. The 

 extreme lateral portions fill and then the whole canals. As the uterus 

 fills with eggs these passages expand centrally and the transverse por- 

 tion anteriorly, so that the cavity of the uterus becomes one continuous 

 sac. The extension from this stage is by regular outpocketing no 

 different from that found in Anoplocephala and Bertiella, except that 

 the pockets are of necessity shorter since the uterus does not begin as a 

 simple tube. The development of the uterus continues until nearly all 

 the organs of the proglottid are reabsorbed. The cirrus pouch and 



