526 DAVID BRYCE ON TWO NEW SPECIES OF PHILODINA. 



the dorsal toes arrive and attach themselves behind the others, 

 while the spurs take a position immediately at the rear of all 

 the toes, which are at once invaginated, their tips remaining 

 affixed. By the bending of the foot forwards underneath the 

 body, the dorsally placed toes are brought into use as well as 

 the terminal. The latter pair are stout, of moderate length, 

 and of two joints, the distal being telescopically retractile within 

 the proximal. 



Each pair of toes is capable of independent motion, and must 

 therefore be provided with distinct muscles and controlUng 

 nerves. All four toes are broadly truncate, and their tips are 

 pierced with several pores, from w^hich ducts pass upwards 

 through the foot to the cells which form the foot-glands. These 

 glands are each built up of a series of large nucleate cells 

 extending from a little way above the spurs as far as or, in 

 some cases, even beyond the anus. The mucus secreted by them 

 passes down the ducts, and is exuded through the pores in the 

 tips of the toes. 



In the three-toed species of Rotifer and Callidina the central 

 toe is dorsal, the external toes terminal ; but there is practically 

 no interval between their bases, and their movements are 

 usually simultaneous. 



The two new species whose descriptions are appended 

 resemble each other in several particulars. Both are rather 

 below the usual size of Philodinae, and both usually adopt a 

 squatting position when feeding. In each case the eyes are 

 frequently indefinable, and when visible are small and round. 

 The spurs are so nearly alike that, while they absolutely 

 distinguish these two species from all others described, they do 

 not afford reliable distinction between the one species and the 

 other. Fortunately, however, the two forms are readily recog- 

 nised by the skin, which in P. rugosa is rough and more or less 

 opaque, and in P. nemorcdis is smooth and transparent. 



Philodina nemoealis, n. sp. 



Sp. Ch. — Rather small for the genus. Skin smooth. Dental 

 formula, 2/2. Foot of four joints; spurs short, slender, acute, 

 and separated by an interstice nearly as w^ide as their length. 



