110 NYCTAGINACEAE 



margin ; peduncles slender, 2-7 cm. long, usually densely viscid-villous ; involucral bracts lance- 

 olate to linear-lanceolate, 7-9 mm. long; perianth 9-14 mm. long, densely viscid-villous, rose- 

 colored; fruit 4-8 mm. long, wings usually 3, truncate or acutely prolonged above the body of 

 the fruit, narrowed below, body of the fruit indurate, beaked above, coarsely rugose-reticulate, 

 the reticulation extending into the irregular wings. 



In sandy soil, Lower Sonoran Zone; southern Nevada and southeastern California south to western Sonora 

 and Lower California. Type locality: Arizona. March-Oct. 



Abronia pinetorum Abrams and Abronia aurita Abrams, Bull. Torrey Club 32: 537. 1905. Stout, much 

 branched forms, occasionally almost glabrate, characterized by large perianths 2.5-3 cm. long and by fruits 

 larger than those of Abronia villosa. Western and central Riverside County, Orange County, and central San 

 Diego County, California. 



4. Abronia umbellata Lam. Beach Sand-verbena or Abronia. Fig. 1589. 



Abronia umbellata Lam. Tab. Encyc. 1: 469. 1791. 

 Tricratus admirabilis L'Her. ex Willd. Sp. PI. 1: 807. 1797. 

 Abronia californica Raeusch. Nom. Bot. ed. 3. 191. 1797. 

 Abronia rotundiflora Gaertner, Fruct. 3: 181. 1807. 

 Abronia alba Eastw. Proc. Calif. Acad. III. 1: 97. 1898. 



Perennial prostrate herbs, branches mostly slender, 2-10 dm. long, nodes long or short de- 

 pending on growing conditions, minutely viscid-puberulent to viscid-pubescent. Leaves oval or 

 rhombic-oval to lance-elliptic or lance-oblong, 2.5-7 cm. long, entire or sinuate in outline, 

 petioles typically as long or longer than the blades ; peduncles 2-12 cm. long, equaling or shorter 

 than the nodes except in vigorous specimens, viscid-puberulent, bracts lanceolate, 4-6 mm. long ; 

 perianth rose, rarely white, tube 10-15 mm. long, glanduiar-pubescent, limb 8-10 mm. broad with 

 deeply emarginate lobes; fruit 7-12 mm. long, about as wide as long, glandular-villous above, 

 becoming glabrate, body of the fruit indurate with a short beak, wings 2-5, usually 4, some- 

 times none, irregularly and faintly reticulate-veined, narrowed below, widened above, truncate 

 or with rounded lobes, or tapering to the beak. 



Sea beaches, Upper Sonoran, Humid Transition and Canadian Zones; British Columbia south to Lower 

 California. Type locality: Monterey, California. Jan.-Dec. 



The form characterized by having the wings of the fruit narrowed above as well as below is much more 

 commonly found in the northern part of the range of the species. In the southern extension, from the type 

 locality southward, the form having the wings of the fruit produced upward in rounded lobes is more prevalent. 

 At Monterey both types are common. 



Abronia acutalata, A. breviflora, A. minor, A. neurophylla, A. insularis, A, variabilis Standley (Contr. U.S. 

 Nat. Herb. 12: 311-314. 1909), are all minor variations. 



5. Abronia latifolia Eschsch. Yellow Sand-verbena or Abronia. Fig. 1590. 



Abronia latifolia Eschsch. Mem. Acad. St.-Petersb. 5: 271. 1826. 

 Abronia arenaria Menzies ex Hook. Exot. Fl. 3: pi. 193. 1827. 

 Tricanthus arenarius Spreng. Syst. Cur. Post. 53. 1827. 



Prostrate glandular perennial, with thick fleshy roots and stout stems, 2-10 dm. long. Leaves 

 thick, orbicular to reniform, rarely rounded-deltoid, 3-6 cm. long; petioles 1.5-4 cm. long, 

 densely viscid-puberulent ; blades 1-3 cm. long, glabrate ; peduncles 2-4 cm. long ; bracts 5-6, 

 ovate to ovate-lanceolate, 6-8 mm. long ; perianth 13-18 mm. long, the tube slender, glandular- 

 villous, greenish yellow, the limb 5-8 mm. broad, bright yellow, lobes shallowly emarginate; 

 fruit 8—15 mm. long, coriaceous, puberulent, mostly biturbinate with 5 wing-like reticulate- 

 veined lobes, these attenuate upward on the body of the fruit. 



On sea beaches, Canadian, Humid Transition and Upper Sonoran Zones; British Columbia to Santa Barbara 

 County, California. Type locality: California. Feb.-Nov. 



6. Abronia maritima Nutt. Red Sand-verbena or Abronia. Fig. 1591. 



Abronia maritima Nutt. ex S. Wats. Bot. Calif. 2: 4. 1880. 



Prostrate perennials, glandular-puberulent to villous throughout with fleshy roots and very 

 stout trailing stems, 2-10 dm. long. Leaves thick, 4-6 cm. long, petioles 1-2 cm., blades thick, 

 3-6 cm. long, oval or oval-oblong, cuneate or rounded at base, rounded at apex ; peduncles 3-8 cm. 

 long, bracts lanceolate to ovate-lanceolate, 6-8 mm. long, perianth 11-14 mm. long, dark crim- 

 son, limb 3-5 mm. broad; fruit coriaceous, densely puberulent above and glabrate below, 10—14 

 mm. long, turbinate, wing-like lobes irregular, usually 5, truncate above, scarcely extending 

 above the body of the fruit, attenuate below, body and base of the fruit coarsely reticulate- 

 veined. 



On sea beaches. Upper and Lower Sonoran Zones; San Luis Obispo County south to Lower California and 

 Sinaloa, Mexico. Type locality: San Pedro, California. Feb.-Nov. 



7. Abronia nana var. Covillei (Heimerl) Munz. Coville's Dwarf Abronia. 



Fig. 1592. 



Abronia Covillei Heimerl, Smiths. Misc. Coll. 52: 197. 1908. 

 Abronia nana var. Covillei Munz, Man. S. Calif. 150, 598. 1935. 



Densely cespitose, glandular perennials with woody roots and short, stout branching caudex, 

 puberulent throughout with short, scattered hairs. Leaves clustered, 2-4 cm. long, petioles 

 slender, 1-3 cm. long, blades ovate to oblong, 5-15 mm. long; peduncles scape-like, 4-10 cm. 

 long; bracts 4-6, 4-6 mm. long, lanceolate to narrowly ovate; perianth 10-13 mm. long, white 

 with a pink tube ; fruit 6-8 mm. long, obcordate with 5 thin-walled, regular, wing-like lobes. 



Arid Transition Zone; Inyo Mountains, Inyo County, California, and adjacent ranges in Nevada south to 

 the San Bernardino Mountains, California. Type locality: Inyo Mountains, California. June-Aug. 



