270 BRASSICACEAE 



ing or horizontal, 6-8 mm. long, often 2 mm. wide, acute at base, obtuse at apex; style minute; 

 seeds less than 1 mm. long, always in 2 rows. 



Usually in sandy soil, Upper and Lower Sonoran Zones; Contra Costa County, California, to southern Cali- 

 fornia, east to the Mojave and Colorado Deserts. Type locality: California. April- June. 



Descurainia pinnata subsp. halictorum (Cockerell) Detling, Amer. Midi. Nat. 22: SOS. 1939. (.Sophia 

 halictorum Cockerell, Bull. Torrey Club 25: 460. 1898.) Characterized by divergent and ascending pedicels 

 rather than horizontal, pale yellow flowers, siliques shorter than those of the species, and terminal leaflets of 

 the upper leaves usually elongated; seeds distinctly in 2 rows. Southeastern Oregon and Idaho and eastern 

 slope of the Sierra Nevada, California, east to western Texas and Oklahoma. 



Descurainia pinnata subsp. glabra (Woot. & Standi.) Detling, Amer. Midi. Nat. 22: 507. 1939. 

 (Sophia glabra Woot. & Standi. Contr. U.S. Nat. Herb. 16: 127. 1913.) inflorescence glabrous, flowers pale 

 yellow, siliques shorter than the preceding subspecies (4-6 mm. long), terminal leaflets rounded not elongate, 

 seeds distinctly in 2 rows. Most abundant form in the Sonoran Desert area. 



Descurainia pinnata subsp. intermedia (Rydb.) Detling, Amer. Midi. Nat. 22: 511. 1939. (Sophia 

 intermedia Rydb. Mem. N.Y. Bot. Gard. 1: 184. 1900.) Plants non-glandular, nearly glabrous; petals yellow, 

 about 2 mm. long, pedicels spreading, shorter than the siliques, seeds more or less distinctly in 2 rows. North- 

 eastern California and adjacent Nevada, and southern Montana to western Wyoming and Colorado. 



Descurainia pinnata subsp. Nelsonii (Rydb.) Detling, Amer. Midi. Nat. 22: 512. 1939. (Sophia Nel- 

 sonii Rydb. Bull. Torrey Club 34: 436. 1907.) Plants non-glandular, nearly glabrous, petals yellow, about 1.5 

 mm. long; pedicels and siliques short, erect; seeds more or less distinctly in 2 rows. Eastern Oregon and 

 Washington, east to Montana, Wyoming, and Colorado. 



Descurainia pinnata subsp. filipes (A. Gray) Detling. Amer. Midi. Nat. 22: 513. 1939. (Sisymbrium 

 incisum var. filipes A. Gray, Mem. Amer. Acad. II. 4: 8. 1849.) Plants non-glandular, nearly glabrous; 

 petals yellow, 2 mm. long; pedicels longer than the siliques, the siliques spreading or erect. Eastern slopes of 

 Cascade Mountains, Washington and Oregon, and south in the Sierra Nevada, California, to Yosemite Valley. 

 Also in the northern Rocky Mountain region. 



6. Descurainia paradisa (A. Nels. & Kennedy) O. E. Schulz.. Nevada 



Tansy-mustard. Fig. 2006. 



Sophia paradisa A. Nels. & Kennedy, Proc. Biol. Soc. Wash. 19: 155. 1906. 

 Descurainia paradisa O. E. Schulz, Pflanzenreich 4 105 : 331. 1924. 



Gray, canescent annual herbs, branching from the base, 1-2.5 dm. high. Leaves bipinnate 

 or bipinnatifid into short, mostly linear-oblong lobes ; inflorescence scantily pubescent, glandu- 

 lar ; flowers whitish-yellow ; pedicels 3-4 mm. long, ascending, sometimes curved ; pods 1 . 5-3 . 5 

 mm. long, on same axis as pedicel or nearly erect, elliptic in outline, obtuse at the apex ; style 

 minute ; seeds less than 1 mm. long, distinctly in 2 rows. 



Upper Sonoran Zone; Great Basin region from southeastern Oregon to Inyo County, California. Type 

 locality: Paradise Valley, Humboldt County, Nevada. Closely related to the preceding species complex. 

 April-May. 



18. CAKILE [Tourn.] Mill. Gard. Diet. Abr. ed. 4. 1754. 



Fleshy glabrous branching annual herbs, with purple or white flowers. Style none. 

 Pods sessile, flattened or ridged, indehiscent, 2-jointed, the joints 1 -celled and 1 -seeded, 

 or the lower seedless, the upper deciduous when ripe, the lower persistent. Cotyledons 

 accumbent. [Old Arabic name.] 



A genus of 4 species, natives of sea and lake shores of north temperate and tropical regions. Type 

 species, Bunias Cakile L. 



Lower joint of silique with two slightly recurved horn-like protuberances at apex; leaves pinnatifid. 



1. C. maritima. 

 Lower joint of silique without expanding horn-like processes; leaves merely sinuate-dentate. 



2. C. edentula califomica. 



1. Cakile maritima Scop. Sea Rocket. Fig. 2007. 



Bunias Cakile L. Sp. PI. 670. 1753. 



Cakile maritima Scop. Fl. Cam. ed. 2. 2: 35. 1772. 



Plants fleshy and glabrous, branching from the base, the branches procumbent or decum- 

 bent, often 4-5 dm. long. Leaves deeply pinnatifid, 4-8 cm. long, the lobes varying from 4-8, 

 oblong or linear, 5-15 mm. long, rounded at apex; sepals 3 mm. long; petals rose-colored, 8-10 

 mm. long; fruiting racemes 20-35 cm. long; pedicels stout, about 2 mm. long; pods about 15 mm. 

 long; the lower joint half the length of the upper, bearing 2 triangular protuberances at the 

 apex, upper joint flattened, entire at the apex. 



Beach sand; along the coast from Sonoma County to San Mateo County, California. Native of Europe and 

 the Mediterranean region. June-Nov. 



2. Cakile edentula var. califomica (Heller) Fernald. California Sea 



Rocket. Fig. 2008. 



Cakile califomica Heller, Muhlenbergia 3: 10. 1907. 



Cakile edentula var. califomica Fernald, Rhodora 24: 23. 1922. 



Plants fleshy, the main stem stout, erect, branching from the base, the lower lateral branches 

 often decumbent. Leaves 15-40 mm. long, oblanceolate, rounded or obtuse at the apex, nar- 

 rowed to a stout petiole, sinuate-dentate ; racemes becoming 1-2 dm. long in fruit ; pedicels stout, 

 3-5 mm. long; sepals 3-4 mm. long; petals purple-tinged, 6 mm. long; pods 15 mm. long, upper 

 joint broader than the lower, flattened and angled, narrowed to a stout retuse beak. 



Beach sands; the typical species ranges from Labrador to Florida, the variety lacustris Fernald is con- 

 fined to the Great Lakes, and the variety califomica extends along the shores of the Pacific from British 

 Columbia to southern California. May-Nov. 



