MUSTARD FAMILY 283 



3. Dentaria pachystigma S. Wats. Stout-beaked Toothwort. Fig. 2042. 



Dentaria californica var. pachystigma S. Wats. Proc. Amer. Acad. 14: 289. 1879. 

 Dentaria pachystigma S. Wats, in A. Gray, Syn. Fl. N. Amer. I 1 : 155. 1895. 



Glabrous, the stems stout, about IS cm. high. Rhizomal leaves cordate-orbicular, crenately 

 toothed, stem leaves 2 or 3, approximate, simple, cordate or reniform, crenate or sinuate or 

 more frequently coarsely dentate ; raceme nearly sessile ; pods 2 . 5-5 cm. long, 3-4 mm. broad ; 

 style short and stout; seeds nearly orbicular; cotyledons thick. 



The typical form of this species is little known. It was originally collected in the mountains of Plumas 

 County, California, and our illustration is made from the type specimens in the Gray Herbarium, Harvard 

 University. 



Dentaria pachystigma var. corymbosa (Jepson) Abrams. (Dentaria corymbosa Jepson, Man. Fl. PI. 

 Calif 426 1925.) Style narrower, scarcely flattened, 4-10 mm. long; racemes short, almost corymbose. This 

 is the common form of the species, ranging from Shasta to the South Yollo Bolly Mountains, Lake County, 

 and to Tulare County in the Sierra Nevada, California. 



Dentaria pachystigma var. dissectifolia Detling, Amer. Journ. Bot. 23: 575. 1936. Stem leaves pin- 

 nately 3-5-foliolate. Serpentine rocks, Mogalia, Butte County, California. 



4. Dentaria californica Nutt. California Toothwort. Fig. 2043. 



Dentaria californica Nutt. in Torr. & Gray, Fl. N. Amer. 1: 88. 1838. 

 Cardaminc californica Greene, Fl. Fran. 266. 1891. 



Glabrous or slightly pubescent, often more or less purplish. Leaves rather thin, not fleshy ; 

 rhizomal leaves entire or trifoliolate, mostly orbicular, the blades or leaflets commonly dentate 

 or lobed; stem leaves 3-foliolate, the leaflets lanceolate-oblong to nearly linear, more or less 

 toothed; flowers white or pale rose; pods 2.5-6 cm. long, 2-3 mm. wide; styles 3-5 mm. long. 



Shady banks and rich woods, Upper Sonoran and Transition Zones; southern Oregon to northern Lower 

 California. Type locality: Monterey, California. Feb.-April. 



Dentaria californica var. cuneata (Greene) Detling, Amer. Journ. Bot. 23: 576. 1936. (Cardamine 

 cuneata Greene, Bull. Calif. Acad. 1:74. 1888. Cardamine californica subsp. cuneata O. h. Schulz, Hot. 

 Tahrb. 32: 386. 1903.) Leaves more ample and at least those of the stem 5-foliolate. Santa Lucia and Gabilan 

 Mountains, Monterey County, California. Type locality: near Jolon. 



Dentaria californica var. sinuata (Greene) Detling, Amer. Journ. Bot. 23: 576. 1936. (Dentaria sinu- 

 ata Greene, Pittonia 3: 123. 1896. Cardamine californica var. sinuata O. E. Schulz, Bot. Jahrb. 32: 387. 

 1903 Dentaria californica var. Tracvi Jepson, Man. Fl. PI. Calif. 426. 1925.) Plants 15-35 cm. high. Basal 

 leaf simple, round-reniform, cordate at base, 5-7.5 cm. broad, sinuately lobed; stem leaves 2 or 3, 3-5-foho- 

 late lobed or coarsely toothed; petals 10-15 mm. long, rose-purple; style 3-5 mm. long. This variety grows 

 on wooded slopes in the Humid Transition Zone from Coos County, Oregon, to Mendocino County, California. 

 Type locality: Redwoods, near Crescent City, California. 



Dentaria californica var. integrifolia Detling, Amer. Journ. Bot. 23: 576. 1936. (Dentaria integrifolia 

 Nutt. in Torr. & Gray, Fl. N. Amer. 1: 88. 1838.) Glabrous or slightly pubescent, stem 1.5-6 dm._ high, stout. 

 Leaves rather thick and fleshy, the blades or leaflets entire; basal leaves simple or trifoliolate, orbicular, ovate 

 or reniform 2-8 cm. long; stem leaves 3-foliolate or sometimes 5-foliolate, the leaflets ovate to lanceolate or linear; 

 flowers white; petals 10-15 mm. long, scarcely spreading; pods flattened, 25-35 mm. long; style stout, 3-4 mm. 

 long. Low moist fields near the coast, Sonoran and Transition Zones; central California. Type locality: 

 "Plains of Monterey, California." 



Dentaria californica var. cardiophylla (Greene) Detling, Amer. Jour. Bot. 23: 576. 1936. (Cardamine 

 cardiophylla Greene, Fl. Fran. 266. 1891. Dentaria cardiophylla Robinson, Syn. Fl. N. Amer. I 1 : ls5. 1895. 

 Dentaria integrifolia var. cardiophylla Jepson, Man. Fl. PI. Calif. 426. 1925.) Glabrous, the stems usually 

 simple from a small tuber, 1.5-3 dm. high. Leaves all simple, the basal round-cordate, those of the stem 

 round-cordate to lanceolate, sinuate to acutely toothed, 1.5-3 cm. long, petals rose-colored; pods 25-35 mm. 

 long, 2 mm. broad; style slender, 3-4 mm. long. Shady banks, Sonoran and Transition Zones; Inner North 

 Coast Ranges and northern Sierra Nevada, California. Type locality: Vaca Mountains, Solano County, Cali- 

 fornia. 



29. IDAHOA A. Nels. & Macbr. Bot. Gaz. 56:474. 1913. 



Low slender scapose glabrous annuals with lyrate basal leaves. Flowers solitary. 

 Sepals broad, erect. Petals white, small, spatulate. Silicles suborbicular, strongly flat- 

 tened parallel to the partition. Seeds in 2 rows in each cell, reticulate, broadly winged. 

 [Named for the state of Idaho.] 



A monotypic genus of the Great Basin region. 



1. Idahoa scapigera (Hook.) A. Nels. & Macbr. Flat-pod. Fig. 2044. 



Platyspermum scapigerum Hook. Fl. Bor. Amer. 1: 68. pi. 8. fig. B. 1830. 

 Idahoa scapigera A. Nels. & Macbr. Bot. Gaz. 56: 474. 1913. 



Leaves rosulate, petioled, ovate, lyrate or sometimes entire; scapes several, 2-15 cm. high; 

 sepals 2 mm. long, nearly equaling the spatulate petals ; pod suborbicular, 6-10 mm. long, 8-12- 

 seeded ; seeds broadly winged, 5 mm. broad. 



Mountain valleys in moist gravelly soils, Arid Transition Zone; eastern Washington and Idaho to north- 

 eastern Nevada and Lassen and Santa Clara Counties, California. Type locality: moist rocks and in shallow 

 soils, at the Celilo Falls, Columbia River. March-April. 



30. TROPIDOCARPUM Hook. Ic. PI. 1 : pi. 43. 1836. 



Slender, branching annuals, more or less hirsute with simple and forked hairs inter- 

 mingling. Leaves pinnatifid ; flowers small, in loose, leafy racemes. Sepals oblong, con- 

 cave, equal at base, spreading. Petals yellow, spatulate-obovate. Stamens tetradynamous ; 



