180 RANUNCULACEAE 



late, 10-18 mm. long, exceeding the spur; lamina of the petals cream-colored, 5-8 mm. long; 

 spur shorter than the sepals, usually incurved at the apex ; follicles about 2 cm. long, pubescent. 

 Open woods, Arid Transition and Canadian Zones; British Columbia to eastern Oregon, east to Alberta, 

 Wyoming, and Utah. Type locality: "Wasatch and Uintah Mountains," Utah. June- Aug. 



2. Aquilegia formdsa Fischer. Northwest Crimson Columbine. Fig. 1775. 



Aquilegia formosa Fischer in DC. Prod. 1 : SO. 1824. 



Aquilegia canadensis var. fortnosa S. Wats. Bot. King Expl. 10. 1871. 



Aquilegia columbiana Rydb. Bull. Torrey Club 29: 145. 1902. 



Stems 5-10 dm. high, glabrous or sparingly pubescent. Leaves biternate; leaflets 2-4 cm. 

 long, 2-3-cleft, the segments crenately lobed or toothed at the apex; sepals and spurs crimson; 

 sepals ovate-lanceolate, acute or acuminate, about 2 cm. long; lamina of petals 6-10 mm. long, 

 truncate or rounded, yellow; spur 12-18 mm. long, straight; follicles densely pubescent, erect, 

 20-30 mm. long. 



Open woods, Transition and Canadian Zones; Alaska to northern and eastern California, Nevada, Utah, 

 and Montana. Type locality: "In Kamchatka." June- Aug. 



Aquilegia formosa var. wawawensis (Payson) St. John, Research Stud. St. Coll. Wash. 1:97. 1929. 

 {A. wawawensis Payson, Contr. U.S. Nat. Herb. 20: 14S. 1918.) This is a form differing chiefly in the 

 thinner leaves and light crimson or scarlet sepals and spurs. Along the banks of the Snake River, near Wawa- 

 wai, Washington. 



Aquilegia formosa var. truncata (Fisch. & Mey.) M. E. Jones, Zoe 4: 2S9. 1893. {Aquilegia truncata 

 Fisch. & Mey. Ind. Sem. Hort. Petrop. 9: Suppl. 8. 1843; A. calif ornica Lindl. Gard. Chron. 1854:836. 

 1854; A. hypolasia Greene, Leaflets Bot. Obs. 2: 141. 1911; A. adiantoides Greene, op. cit.; A. emarginata 

 Eastw. Leaflets West. Bot. 2: 7. 1937.) Closely resembling the species in general habit. Sepals and spurs 

 scarlet, tinged with yellow; orifice of spur almost truncate, the lamina 1-3 mm. long; follicles 15-20 mm. long. 

 Open woods and shady banks, Upper Sonoran to Canadian Zones; southern Oregon to northern Lower Cali- 

 foria and western Nevada. Type locality: Fort Ross, California. 



Aquilegia formosa var. pauciflora (Greene) Payson, Contr. LT.S. Nat. Herb. 20: 144. 1918. (A. Pauci- 

 flora Greene, Leaflets Bot. Obs. 1: 76. 1904.) Leaves mostly basal and tufted; flowering stem scape-like, 2-3 

 dm. high, few-flowered; lamina of petal 2-3 mm. long. This is an alpine form of var. truncata inhabiting the 

 higher altitudes of the Canadian and Hudsonian Zones of the Sierra Nevada, California. 



Aquilegia mohavensis Munz, Leaflets West. Bot. 2: 66. 1938. This recently described species is closely 

 related to A. formosa and probably only a variety of that polymorphic species. Leaves glabrous, nearly triter- 

 nate, the leaflets being almost separate and small, 1.5-2.5 cm. long; sepals 7-8 mm. long, spreading, greenish 

 yellow margined with red; spurs of the petals yellowish red, 13-15 mm. long, blade 3 mm. long, round-truncate; 

 follicles about 15 mm. long. Desert ranges (New York Mountains), eastern Mojave Desert, California. 



3. Aquilegia eximia Van Houtte. Van Houtte's Columbine. Fig. 1776. 



Aquilegia eximia Van Houtte ex Planch, Fl. Serres II. 2: 15. 1857. 

 Aquilegia Tracyi Jepson, Fl. W. Mid. Calif, ed. 2. 165. 1911. 



Plants glandular-puberulent throughout, the stems 5-10 dm. high. Basal leaves triternate; 

 leaflets 2-4 cm. long, more or less glandular-pubescent, their teeth inclined to be obtuse instead 

 of rounded; sepals about 15 mm. long, ovate-lanceolate, reflexed, scarlet; spurs scarlet below, 

 tinged with yellow toward the orifice, 20-25 mm. long, 8-10 mm. broad at the orifice and sloping 

 backward; lamina obsolete; follicles 15-20 mm. long. 



Moist rocky banks, Upper Sonoran and Humid Transition Zones; California Coast Ranges from Mendo- 

 cino County to San Mateo County. Type locality: not stated. April-Aug. 



4. Aquilegia pubescens Coville. Coville's Columbine. Fig. 1777. 



Aquilegia pubescens Coville, Contr. U.S. Nat. Herb. 4: 56. pi. 1. 1893. 



Stems 2-3 dm. high, minutely pubescent throughout or glabrous below. Basal leaves ternate 

 or biternate; leaflets 3-4 cm. long, with rounded lobes, glabrate to densely pubescent; stem 

 leaves few; flowers erect, yellow throughout; sepals oblong, abruptly acute, 20-25 mm. long, 

 spreading horizontally; spurs slender, straight, 3.5-4 cm. long; lamina 8-14 mm. long; follicles 

 about 2 cm. long, pubescent. 



Moist rocky soils, Boreal Zones; Sierra Nevada, mostly above 9,000 feet altitude, Mariposa County to 

 Tulare County, California. Type locality: Mineral King, Tulare County, California. June-Aug. 



9. DELPHINIUM L. Sp. PI. 530. 1753. 



Annual or perennial herbs with erect simple or branched stems, palmately lobed or 

 divided leaves and showy irregular flowers. Sepals 5, petaloid, the upper one prolonged 

 into a spur. Petals 2 or 4, the two upper spurred, the other two when present lateral and 

 small. Pistils 1 or 3, sessile, many-ovuled, forming follicles in fruit. 



A genus of about 126 species inhabiting the north temperate regions. Type species, Delphinium Con- 

 solida L. 



Annual; pistil 1. \. D. Ajacis. 



Perennials; pistils 3. 



Flowers red or yellow. 



Flowers red or rose-colored. 



Leaves divided into narrowly linear or lanceolate divisions; herbage and calyx puberulent. 



2. D. cardinale. 

 Leaves divided into broad primary divisions. 



Flowers rose-colored, often tinged with purple in drying; sepals widely spreading or reflexed. 



3. D. Purpusii. 



Flowers red; sepals only slightly spreading, never reflexed. 4. D. nudicaule. 



