352 SAXIFRAGACEAE 



a shining minutely punctate testa. [Dedicated to Dr. Boykin, a physician from Georgia, 

 a contemporary of Nuttall.] 



A North American and eastern Asiatic genus of about 8 species. Type species, Boykinia aconitifolia Nutt. 



Leaves cleft or incised, with acute teeth; petals conspicuously exceeding the sepals. 



Stipules, at least the upper, foliaceous, often large. 1. B. major. 



Stipules small, brown-scarious, mostly reduced to brownish bristles. 2. B. elata. 



Leaves merely crenately lobed, the lobes crenate-serrate with broad mucronate teeth; petals scarcely exceeding 

 the sepals. 3. B. rotundifolia. 



1. Boykinia major A. Gray. Mountain Boykinia. Fig. 2228. 



Boykinia occidentalis var. elata A. Gray, Proc. Amer. Acad. 8: 383, in part. 1872. 

 Boykinia major A. Gray, Bot. Calif. 1 : 196. 1876. 

 Therofon major Kuntze, Rev. Gen. PI. 1 : 227. 1891. 

 Therofon majus Wheelock, Bull. Torrey Club 23: 70. 1896. 

 Boykinia major var. intermedia Piper, Erythea 7: 172. 1899. 



Tall and stout; stem 3-10 dm. high, more or less glandular-villous with brown hairs. 

 Basal and lower cauline leaves long-petioled, with scarious stipules ; petioles 1-2 dm. long, also 

 glandular-villous ; blades renif orm or rounded-cordate in outline, glabrous except on the veins 

 beneath, 5-20 cm. wide, 5-7-cleft from a third to two-thirds of the way down, again cleft and 

 coarsely serrate with lance-ovate gland-tipped teeth ; middle cauline leaves _ similar but with 

 shorter petioles and large foliaceous stipules ; the uppermost ovate and sessile, the foliaceous 

 stipules and leaf-blades often connate ; inflorescence a relatively dense, many-flowered, cymoid 

 panicle, elongated in fruit, densely glandular-puberulent ; hypanthium broadly turbinate-cam- 

 panulate, becoming urceolate in fruit, without the sepals 2-3 mm. long, glandular-puberulent ; 

 sepals triangular-lanceolate, 3 mm. long; petals white, broadly oval or obovate, clawed, 5-7 

 mm. long. 



Along mountain streams and in damp woods, Humid and Arid Transition Zones; Montana westward to 

 Washington and southward to Trinity and Madera Counties, California. Type locality: Sierra Nevada, Cali- 

 fornia. June-Sept. 



2. Boykinia elata (Nutt.) Greene. Brook Foam. Coast Boykinia. Fig. 2229. 



Saxifraga elata Nutt. in Torr. & Gray, Fl. N. Amer. 1: 575. 1840. 



Boykinia occidentalis Torr. & Gray, Fl. Amer. 1: 577. 1840. 



Boykinia occidentalis var. elata A. Gray, Proc. Amer. Acad. 8: 383. 1872. 



Boykinia elata Greene, Fl. Fran. 190. 1891. 



Therofon occidentale Kuntze, Rev. Gen. PI. 1: 227. 1891. 



Therofon elatum Greene, Man. Bay Reg. 121. 1894. 



Boykinia Nuttallii Macoun, Can. Rec. Sci. 6: 408. 1895. 



Therophon circinnatum Rosend. & Rydb. N. Amer. Fl. 22: 124. 1905. 



Erect and commonly slender; stems sometimes more than one from a branched rootstock, 

 2-6 dm. high, more or less brown-hairy to glabrate below, densely glandular-puberulent above. 

 Basal leaves thin, glabrous or sometimes sparsely covered beneath by distinct rusty hairs, 2-8 cm. 

 wide, reniform to ovate-cordate, usually shallowly to relatively deeply 5-7-lobed, the lobes serrate 

 with broadly ovate bristle-pointed teeth; their petioles 5-15 cm. long, sparingly pilose with 

 brownish hairs, the stipules brown-scarious, small, the greater portion reduced to several rusty 

 bristles ; cauline leaves similar, though successively smaller and with shorter petioles, becoming 

 cuneate at the base, the uppermost rhombic in outline; inflorescence a many-flowered, bracteate 

 panicle, more usually consisting of secund racemes, densely glandular-puberulent; bractlets 

 usually foliaceous, spatulate, incised; hypanthium campanulate, 2-3 mm. long, glandular below, 

 often purplish, in fruit becoming urceolate and 3-5 mm. long ; sepals lanceolate-acuminate, gla- 

 brous, 1.5-2 mm. long; petals white, often pinkish in age, mostly oblanceolate, slightly narrowed 

 to a broad claw, 3-4 mm. long. 



Along streams and in damp woods, Humid and Arid Transition Zones; from British Columbia southward 

 in the coastal mountains to Los Angeles County, and inland west of the Cascades to the central Sierra Nevada, 

 California. Type locality : Chinook Point, mouth of the Columbia River. May- July. 



3. Boykinia rotundifolia Parry. Round-leaved Boykinia. Fig. 2230. 



Boykinia rotundifolia Parry ex A. Gray, Proc. Amer. Acad. 13: 371. 1878. 

 Therophon rotundifolium Wheelock, Bull. Torrey Club 23: 70. 1896. 



Very tall and stout ; stems 5-10 dm. high, densely glandular-villous. Basal and lower cau- 

 line leaves long-petioled, rounded-reniform to almost deltoid, 8-15 cm. wide, with numerous shal- 

 low, rounded lobes, glabrate above except on the veins, usually puberulent beneath, the veins more 

 or less hirsutulous ; teeth of the lobes crenate-serrate, apiculate ; petioles 1-2 dm. long, glan- 

 dular-villous, the slightly dilated base with brown bristly hairs; upper cauline leaves 3-5 cm. 

 long, short-petioled, orbicular to rhombic in outline : peduncles mostly in the axils of the 

 reduced upper leaves; flowers short-pedicelled, in fruit drooping and secund on the elongated 

 branches ; hypanthium campanulate-urceolate, 3-4 mm. long, strongly striate, densely glandular- 

 pubescent; sepals ovate-lanceolate, 2 mm. long: petals slightly irregular, obovate-spatulate, 

 3 mm. long, the claw as long as to slightly longer than the blade. 



Banks of streams in canyons, Upper Sonoran and Arid Transition Zones; San Gabriel and San Bernardino 

 Mountains, near Elsinore and in the Cuyama VMley in Santa Barbara County, southern California. Type 

 locality: "Along water-courses, San Bernardino Mountains," California. May-July. 



