354 SAXIFRAGACEAE 



6. JEPSONIA Small, Bull. Torrey Club 23: 18. 1896. 



Perennial acaulescent herbs with tunicated corm-like rootstocks supporting an often 

 elongated caudex and several slender scapes. Leaves alternate, all basal, reniform in out- 

 line, shallowly lobed and toothed, petioled. Flowers in terminal cymes. Hypanthium 

 campanulate, strongly purplish-striate, truncate or acutish at the base, entirely free from 

 the ovary except at the very base. Petals 5, spatulate to oval-elliptic in outline, narrowed 

 to a claw, withering-persistent, adnate to the hypanthium. Stamens 10, shorter than the 

 sepals, the point of attachment of one set immediately below that of the petals ; filaments 

 filiform, the bases adnate to the hypanthium. Ovary superior, the two carpels united to 

 about the middle or higher. Carpels veiny, filling and ultimately protruding beyond the 

 hypanthium, the beaks strongly divergent. [Named for Willis Linn Jepson, Professor of 

 Botany at the University of California and authority on California flora.] 



A genus of limited distribution, confined to California, the islands off its southern coast and northern Lower 

 California. It consists of the following three species. Type species, Jepsonia Parryi (Torr.) Small. 



Sepals shorter than the tube of the hypanthium, the latter at anthesis 3-4 mm. long; flowering scapes autumnal, 

 appearing before the leaves which are vernal. 

 Hypanthium truncate at the base, narrowly campanulate. 1. /. Parryi. 



Hypanthium acutish at the base, open-campanulate. 2. J. heterandra. 



Sepals as long as the tube of the hypanthium or slightly longer, the latter at anthesis 2-2.5 mm. long; flowering 

 scapes appearing at the same time as the basal leaves. 3. /. malvaefoha. 



1. Jepsonia Parryi (Torr.) Small. Coast Jepsonia. Fig. 2231. 



Saxifraga Parryi Torr. Bot. Mex. Bound. 69. 1859. 

 Jepsonia Parryi Small, Bull. Torrey Club 23: 18. 1896. 



Caudex clothed with the firm persistent leaf-bases. Leaves vernal, erect, reniform, dis- 

 tinctly broader than long, sparingly glandular-puberulent, hirsutulous on the veins, 2-8 cm. 

 broad, shallowly lobed and crenately toothed, the teeth apiculate, shallowly to deeply cordate 

 at the base, the sinus usually wide and open; petioles hirsute to glabrate, widened toward the 

 base, 2-6 cm. long; scapes autumnal, often tufted, wiry, 1-3 dm. tall, glandular-puberulent to 

 glabrate in age ; hypanthium narrowly campanulate, strongly purple-striate, truncate at the 

 base, including the short ovate sepals 6 mm. long; petals obovate to oval, narrowed to short 

 claws, lightly purplish-veined, 4-6 mm. long; fruiting carpels flat at the base, protruding 

 beyond the hypanthium, about 7 mm. long, dimorphous, the reniform stigmas subsessile on the 

 broadish beaks in one form, raised on elongated style-like beaks in the other. 



Dry slopes, Lower and Upper Sonoran Zones; mountains of southwestern San Bernardino County southward 

 to San Diego County, California, and adjacent Lower California. Type locality: dry hills near San Diego and 

 San Luis Rey, California. Nov.-Dec. 



2. Jepsonia heterandra Eastw. Foothill Jepsonia. Fig. 2232. 



Jepsonia heterandra Eastw. Bull. Torrey Club 32: 201. 1905. 

 Jepsonia Parryi var. heterandra Jepson, Man. Fl. PI. Calif. 457. 1925. 



Caudex often thick and woody, scaly. Leaves vernal, erect or ascending, suborbicular to 

 rounded-reniform, 3.5-6 cm. broad, shallowly lobed and toothed, more or less pubescent about 

 the veins beneath, cordate at the base, the basal lobes sometimes overlapping; petioles usually 

 hirsute, 2.5-5 cm. long; scapes autumnal, often tufted, 1-3 dm. high, puberulent to glabrate 

 in age ; hypanthium ultimately scarious, open-campanulate, acutish at the base, purplish-striate, 

 including the broadly ovate to triangular-acute sepals 4-5 mm. long, the latter 1-2 mm. long ; 

 petals 5-6 mm. long, ovate to oval, narrowed to short claws ; anthers oblong and acutish at 

 the apex to orbicular ; filaments of varying lengths on the same flower ; carpels strongly reddish- 

 striate, in fruit long-exserted beyond the hypanthium, 5-6 mm. long, the strongly divergent 

 beaks dimorphous, very slender and almost filiform, and slightly shorter than the bodies in 

 one form, much shorter and broader, 1 mm. or less long in the other. 



Dry rocky outcrops, Upper Sonoran Zone; lower Sierra Nevada foothills, Eldorado County to Mariposa 

 County, California. Type locality: banks of the Merced River, near Benton's Mills (Bagby), Mariposa County, 

 California. Aug.-Dec. 



3. Jepsonia malvaefolia (Greene) Small. Island Jepsonia. Fig. 2233. 



Saxifraga malvaefolia Greene, Bull. Torrey Club 9: 121. 1882. 

 Jepsonia malvaefolia Small, Bull. Torrey Club 23: 19. 1896. 

 Jepsonia neonuttalliana Millsp. Field Mus. Bot. Ser. 5: 124. 1923. 



Aerial parts annual from a stout corm-like rootstock, the habit essentially that of the two 

 preceding; younger parts glandular-puberulent. Leaves appearing with the scapes after the first 

 rains, ovate-orbicular to orbicular-reniform, shallowly cordate at the base, 2-3.5 cm. wide, 

 hirsutulous on the veins beneath, irregularly and more sparsely hirsutulous above, closely serrate- 

 dentate toward the base, the shallow crenate obscure lobes irregularly and closely serrate-dentate ; 

 petioles densely hirsutulous, more or less channelled, narrowly scarious-dilated at the base, 2.5-4 

 cm. long ; scapes very slender, 8-20 cm. tall ; inflorescence at first a subcongested cymose panicle ; 

 hypanthium strongly striate, widely campanulate, mostly truncate or sometimes acutish at the 

 base, the narrowly oblong sepals as long as to often slightly longer than the tube ; petals oval- 

 elliptic, conspicuously purplish-veined, about 3.5 mm. long, the claw short and broad; carpels at 

 anthesis united for two-thirds of their lengths, the stigmas subsessile on stout beaks or tipping 



slender style-like beaks. 



Canyon banks and open mesas, Upper Sonoran Zone; Channel Islands (Santa Cruz, Santa Rosa and Santa 

 Catalina Islands) off the coast of southern California, and on the mainland in western San Diego County (vicinity 

 of Hodges Dam), California. Type locality: Santa Rosa Island. Nov.-Dec. 



