418 ROSACEAE 



7. Horkelia Wilderae Parish. Wilder's Horkelia. Fig. 2385. 



Horkelia Wilderae Parish, Bot. Gaz. 38: 460. 1904. 



Potentilla Wilderae Munz & Jtn. Bull. S. Calif. Acad. 24: 8. 1925. 



Potentilla Parryi var. Wilderae Jepson, Man. Fl. PI. Calif. 494. 192S. 



Stems several crowning a deep perpendicular taproot, slender, 2 dm. high ; stipules lanceolate, 

 entire or 1-2-toothed. Leaflets of basal leaves 5-6 pairs, 5-7 cm. long, cuneate, deeply incised, 

 the lobes oblong; cyme diffuse, the flowers numerous on very slender recurved pedicels; hypan- 

 thium glabrate, about 2 mm. broad; sepals lanceolate, 2 mm. long, twice the length of the 

 linear-oblong bractlets ; petals obovate, about equaling the sepals. 



Along trail leading from Barton Flat to South Fork of Santa Ana River, San Bernardino Mountains, Cali- 

 fornia. The only known locality. June— Aug. 



8. Horkelia fusca Lindl. Dusky Horkelia. Fig. 2386.. 



Horkelia fusca Lindl. Bot. Reg. 23: pi. 1997. 1837. 

 Potentilla Douglasii Greene, Pittonia 1 : 103. 1887. 

 Horkelia tenuisecta Rydb. N. Amer. Fl. 22 : 278. 1908. 



Stems from a short woody caudex, 1-3 dm. high, erect and simple, densely glandular above. 

 Leaflets of basal leaves 5-10 pairs, 5-10 mm. long, cuneate or obovate in outline, deeply pin- 

 natifid into linear segments, these sometimes again toothed; cymes corymbose and congested or 

 open ; hypanthium 4 mm. broad, glandular-hirsute ; sepals acuminate, 4 mm. long ; petals cuneate, 

 slightly emarginate, 4 mm. long. 



Meadows and open woods, mainly Transition Zone; Kittitas County, Washington, to the vicinity of Mount 

 Hood, Oregon. Type locality: erroneously attributed to "California." May-Aug. 



Horkelia fusca subsp. capitata (Lindl.) Keck, Lloydia 1: 97. 1938. (Horkelia capitata Lindl. Bot. Reg. 

 23: under pi. 1997. 1837; Potentilla capitata Greene, Pittonia 1: 104. 1887; Horkelia caeruleomontana St. 

 John, Fl. S.E. Wash. 199. 1937.) Stems stout, 4-6 dm. high, glandular-hirsute above. Leaflets of basal leaves 

 5-7 pairs, obovate or oval, 2-3 cm. long, bright green and nearly glabrous, deeply incised-toothed; cymes sub- 

 capitate, purple, subtended by broad palmately divided leaves, equaling or surpassing the head; sepals acuminate, 

 4-5 mm. long, about equaled by the linear bractlets; petals 5-6 mm. long, truncate. Open coniferous forests, 

 Arid Transition and Canadian Zones; Coeur d'Alene Mountains, Idaho, to the Blue and Steens Mountains, 

 eastern Oregon, and Warner Mountains, California. Type locality: "America-boreali-occidentalis," Douglas. 



Horkelia fusca subsp. pseudocapitata (Rydb.) Keck, Lloydia 1: 99. 1938. (Horkelia pseudocapitata 

 Rydb. ex Howell, Fl. N.W. Amer. 1: 180. 1898; H. Brownii Rydb. N. Amer. Fl. 22: 276. 1908; Potentilla 

 Douglasii var. pseudocapitata Jepson, Fl. Calif. 2: 203. 1936.) Leaflets of basal leaves 5-7 pairs, obovate or 

 cuneate, 8-20 cm. long, toothed or incised above; cymes subcapitate, their leaves small, much shorter than the 

 heads; sepals ovate-lanceolate, well exceeding the linear bractlets; petals broadly cuneate, 3-4 mm. long, truncate 

 or slightly emarginate at apex. Border of mountain meadows, Arid Transition and Canadian Zones; Deschutes 

 County, Oregon, to the eastern slopes of the northern Sierra Nevada, California, and adjacent Nevada. Type 

 locality : Janesville, Lassen County, California. 



Horkelia fusca subsp. parviflora (Nutt.) Keck, Lloydia 1: 99. 1938. (Horkelia parviflora Nutt. ex 

 Hook. & Arn. Bot. Beechey 338. 1838; Potentilla Andersonii Greene, Pittonia 1: 104. 1887.) Leaves sparingly 

 pubescent, dark green, decidedly glandular; leaflets 4-8 pairs, cuneate-obovate, cleft at the apex into oblong or 

 lanceolate lobes; cyme capitate or branched; bractlets linear-filiform; sepals ianceolate, 2-3 mm. long; petals 

 broadly cuneate, 2-3 mm. long. Open pine forests, Arid Transition and Canadian Zones; central Idaho and 

 Yellowstone Park to southern Washington and along the Cascades and Sierra Nevada to Tulare County, Cali- 

 fornia. Type locality: "Plains of the Oregon [Columbia] toward the Rocky Mountains." 



Horkelia fusca subsp. filicoides (Crum) Keck, Lloydia 1 : 100. 1938. (Potentilla Douglasii var. fdicoides 

 Crum, Leaflets W. Bot. 1: 100. 1934.) Leaves mostly basal; leaflets deeply pinnatifid into linear segments; 

 cyme openly branched, the flowers solitary or in small glomerules; sepals much exceeding the bractlets. Open 

 pine woods. Transition and Canadian Zones; Cascade Mountains, soutthern Oregon. Type locality: "eight miles 

 north of Diamond Lake, Douglas County, Oregon." 



Horkelia fusca subsp. tenella (S. Wats.) Keck, Lloydia 1: 101. 1938. (Horkelia fusca var. tenella 

 S. Wats. Bot. Calif. 1: 181. 1876; Horkelia tenella Rydb. Bull. Torrey Club 25: 55. 1898.) Leaflets small, 

 palmately divided into linear-oblong segments: cymes dense, usually capitate; otherwise as in suhsp. Parviflora. 

 Dry meadows and flats of volcanic ash, mainly Arid Transition Zone; northern^ Sierra Nevada, Shasta and 

 Tehama Counties to Sierra County, California. Type locality: "Sierra County," California. 



9. Horkelia Clevelandii (Greene) Rydb. Cleveland's Horkelia. Fig. 2387. 



Potentilla Clevelandii Greene, Pittonia 1 : 102. 1887. 

 Horkelia Clevelandii Rydb. Bull. Torrey Club 25: 54. 1898. 

 Horkelia Bolanderi subsp. Clevelandii Keck, Lloydia 1: 94. 1938. 



Stems several from a short caudex, simple, 2-4 dm. high, finely pubescent with spreading 

 hairs, glandular above. Leaflets of basal leaves 7-11 pairs, densely silky-pubescent, broadly 

 obovate-cuneate, 5-10 mm. long and often as broad, deeply crenate-toothed at the apex; cyme 

 with short erect branches ; hypanthium finely pubescent, somewhat glandular ; sepals ovate- 

 lanceolate, acute, 4 mm. long, a little longer than the ovate bractlets ; petals spatulate-oblong, 

 a little longer than the sepals. 



Grassy mountain slopes. Arid Transition Zone; San Jacinto and Cuyamaca Mountains, southern California. 

 Type locality : Laguna, San Diego County, California. June— July. 



10. Horkelia Bolanderi A. Gray. Bolander's Horkelia. Fig. 2388. 



Horkelia Bolanderi A. Gray, Proc. Amer. Acad. 7: 338. 1868. 

 Potentilla Bolanderi Greene, Pittonia 1: 103. 1887. 



Stems several from a slender much branched and somewhat tufted caudex, 1-3 dm. high, 

 finely villous; stipules lanceolate, 5 mm. long, mostly entire. Leaflets of basal leaves 6-11 pairs, 

 loosely to densely villous or hoary-canescent, about 5 mm. long, cuneate, coarsely 3-5-toothed 

 at the apex ; cyme subcapitate or often more open ; hypanthium about 4 mm. broad, saucer- 

 shaped, hoary-pubescent ; bractlets linear-lanceolate, generally much shorter than the broadly 



