ROSE FAMILY 449 



23. SIEVERSIA Willd. Ges. Nat. Freunde Berlin Mag. 5: 397. 1811, 



Perennial usually tufted herbs with rootstocks. Basal leaves lyrately or odd-pinnately 

 divided, usually with smaller segments interspersed ; stem leaves usually reduced ; stipules 

 large, adnate to the petioles. Flowers cymose or solitary, yellow or purple. Hypanthium 

 turbinate or hemispherical, usually 5-bracteolate. Sepals 5. Petals 5. Stamens numerous ; 

 filaments filiform. Pistils numerous; styles terminal, filiform, not jointed, generally 

 elongated in fruit. Fruit a hairy achene ; seed erect, basal. [Name in honor of J. A. C. 

 Sievers, Russian botanist.] 



About IS species of arctic or alpine regions. Type species, Dryas anemonoides Pall. 



Styles much elongated in fruit, plumose to near the apex. 



Petals much exceeding the obtusish sepals; hypanthium turbinate or hemispherical. 1. 5". campanulata. 

 Petals shorter than or about equaling the acute or acuminate sepals. 



Bracts well exceeding the sepals. 2. S. ciliata. 



Bracts mostly shorter than the sepals. 3. 5". canescens. 



Styles not greatly elongated in fruit and not plumose; hypanthium nearly saucer-shaped. 4. S. gracilipes. 



1. Sieversia campanulata (Greene) Rydb. Bell-shaped Purple Avens. Fig. 2474. 



Erythrocoma campanulata Greene, Leaflets Bot. Obs. 1: 178. 1906. 

 Sierversia campanulata Rydb. N. Amer. Fl. 22: 409. 1908. 



Stems 10-25 cm. high from a thick, short rootstock, 1-3-flowered, purple tinged, finely soft- 

 hairy. Basal leaves 3-7 cm. long ; rachis silky-pilose ; principal leaflets rather crowded, obovate- 

 cuneate, 10-15 mm. long, pilose, 3-6-cleft more than half way to the base into linear-oblong 

 divisions ; bractlets linear or linear-subulate, about 5 mm. long ; hypanthium hemispheric, reddish 

 purple, viscid; sepals ovate, obtusish, 6-7 mm. long; petals broadly oval, well exceeding the 

 sepals, tinged and veined with crimson. 



Ridges, Canadian Zone; Olympic Mountains, Washington, and Saddle Mountain, Oregon. Type locality: 

 Olympic Mountains. June— Aug. 



2. Sieversia ciliata (Pursh) G. Don. Long-plumed Purple Avens, Prairie-Smoke. 



Fig. 2475. 



Geum ciliatum Pursh, Fl. Amer. Sept. 352. 1814. 

 Sieversia ciliata G. Don, Gen. Hist. PI. 2: 528. 1832. 

 Geum pubescens Hook. Fl. Bor. Amer. 1: 175. 1832. 

 Erythrocoma ciliata Greene, Leaflets Bot. Obs. 1 : 177. 1906. 



Erect from a stout rootstock, 3-5 dm. high, soft-hairy throughout. Basal leaves tufted, 10- 

 20 cm. long ; principal leaflets 9-19, obovate in outline, 2-5-divided, the divisions cleft and toothed 

 with linear lobes or teeth ; bractlets linear-subulate, usually exceeding the sepals ; sepals ovate- 

 lanceolate, acuminate, 8-10 mm. long ; petals oval, yellowish or tinged with purple, about equaling 

 the sepals; plumose portions of style about 3 cm. long, the upper glabrous portion about Z-A 

 mm. long. 



Open hillsides, Arid Transition and Canadian Zones; British Columbia and Alberta, south to eastern Wash- 

 ington, Utah, and New Mexico. Type locality: banks of the Kooskooskie (Idaho). April-June. 



3. Sieversia canescens (Greene) Rydb. Downy Purple Avens. Fig. 2476. 



Erythrocoma canescens Greene, Leaflets Bot. Obs. 1: 178. 1906. 

 Erythrocoma grisea Greene, Leaflets Bot. Obs. 1: 178. 1906. 

 Sieversia canescens Rydb. N. Amer. Fl. 22 : 409. 1908. 



Stems erect, 2-4 dm. high, finely pilose throughout, mostly 3-flowered. Basal leaves 8-15 cm. 

 long, the rachis hirsute; principal leaflets obovate-cuneate, 10-25 mm. long, densely soft-hairy, 

 ciliate on the margins and veins, mostly 2-3-lobed scarcely half-way to the base, the lobes with 



2 or 3 broadly oblong-ovate teeth ; bractlets oblong-linear or lanceolate, mostly shorter than the 

 sepals; sepals ovate or ovate-lanceolate, acute, about 1 cm. long; petals elliptical, somewhat ex- 

 ceeding the sepals; plumose portion of fruiting style about 3 cm. long, the glabrous portion 



3 mm. long. 



Dry open hillsides, mainly Arid Transition Zone; eastern Oregon and Wyoming, south to California, Arizona, 

 and Chihuahua. Type locality: San Francisco Mountains, Arizona. June-Aug. 



4. Sieversia gracilipes (Piper) Greene. Slender Purple Avens. Fig. 2477. 



Potentilla gracilipes Piper, Bull. Torrey Club 27: 392. 1900. 

 Sieversia gracilipes Greene, Leaflets Bot. Obs. 1: 4. 1903. 

 Acomastylis gracilipes Greene, Leaflets Bot. Obs. 1: 174. 1906. 

 Acomastylis depressa Greene, loc. cit. 



Stems from a thick tufted rootstock and caudex, 6-8 cm. high, pubescent, 1-flowered. Basal 

 leaves many, pinnatifid, silvery sericeous on both surfaces, 4-6 cm. long; divisions 9-21, broadly 

 cuneate in outline, 3-cleft at the apex or entire; stem leaves 1 or 2, small, entire, or 3-lobed; 

 bractlets elliptic, smaller than the sepals; sepals broadly ovate, acutish, 6-7 mm. long; petals 

 orbicular, 7-10 cm. long ; ovaries hairy ; styles glabrous, as long as the fruit. 



Mountain cliffs, Boreal Zones; Blue Mountains, Oregon. Type locality: "face of north cliffs, Blue Moun- 

 tains, Oregon, at the head of Anthony Creek, altitude 8,000 feet." June-Aug. 



