SAXIFRAGE FAMILY 379 



to urceolate, cylindric, hemispheric, turbinate and saucer-shaped, adnate to the lower part 

 of the ovary. Sepals 5, widely ovate to oblong and triangular, erect or spreading, often 

 unequal. Petals small, entire, obovate to spatulate, oblanceolate or linear, usually more 

 or less clawed, sometimes early deciduous or wanting. Stamens 5, opposite the sepals, 

 included or exserted; anthers 2-celled; filaments most often filiform. Ovary approxi- 

 mately half inferior, 1-celled with 2 parietal many-ovuled placentae; capsule-valves nor- 

 mally 2, distinct, usually elongated, capped by the true styles, these sometimes almost or 

 quite wanting. Stigmas inconspicuous to broadly capitate. Capsule opening between and 

 along the more or less divergent beaks. [Named for Johann Heinrich von Heucher, 

 1677-1747, a German professor of medical botany.] 



A genus of about 50 species, natives of North America, from Hudson Bay and Bering Sea to southern Mexico. 

 Type species, Heuchera americana L. 



Stamens equaling or exceeding the sepals (except brevistaminea) ; hypanthium tube longer than the sepals. 

 Hypanthium nearly or quite regular, pale, greenish or whitish, rarely tinged with pink or purple. 



Hypanthium rounded at base and hemispheric, copiously pilose. 1. H. pilosissima. 



Hypanthium acute or acutish at base, turbinate, moderately villous. 

 Panicle becoming diffuse; hypanthium 1.5-3 mm. long. 



Leaves acutely lobed and toothed; seeds with relatively long spine-like processes. 



2. H. glabra. 



Leaves at least the basal with rounded lobes; seeds with the spines very short or obsolete. 



3. H. micrantha. 



Panicle thyrsoid, more or less dense; hypanthium 3.5—5 mm. long. 4. H. maxima. 



Hypanthium oblique at apex, rose-purple; deeply campanulate or urceolate. 



Styles very slender, well-exserted; stamens well-exserted; petals linear or very narrowly oblanceolate, 



less than 0.5 mm. wide, shorter than the stamens. 5. H. rubescens. 



Styles stouter, included; stamens about equaling the sepals or (in brevistaminea) much shorter; petals 

 broadly spatulate, 0.5-1 mm. wide, well exceeding the sepals. 

 Stamens about equaling the sepals. 6. H. elegans. 



Stamens very short, barely reaching the base of the sepals. 7. H. brevistaminea. 



Stamens included, shorter than the sepals; hypanthium pale greenish or yellowish. 

 Hypanthium deeply campanulate to urceolate; panicle spiciform to subcapitate. 

 Flowering branches and petioles hirsute or glandular-pubescent. 

 Flowers yellowish; leaves more or less thick and rigid. 



Leaves glabrous or hairy only on the veins, shining, 3-6 cm. wide, with a sinus at the base. 



8. H. cylindrica. 



Leaves, at least the youngest, more or less densely glandular-puberulent, 1-4 cm. wide, rarely 

 with a basal sinus. 9. H. ovalifolia. 



Flowers greenish; leaves neither thick nor rigid; flowering stems and petioles conspicuously hirsute. 



10. H. chlorantha. 

 Flowering branches and petioles glabrous or merely finely puberulent. 



Flowers yellowish; leaves firm and more or less coriaceous, shining. 11. H. glabella. 



Flowers greenish; leaves thin, neither at all coriaceous nor shining. 12. H. tenuifolia. 



Hypanthium broadly turbinate, 2.5-5 mm. long; panicle narrow but not spiciform; petals elliptic-oblong, 

 acute and mostly acuminate. 13. H. Duranii. 



1. Heuchera pilosissima Fisch. & Mey. Seaside Heuchera. Fig. 2290. 



Heuchera pilosissima Fisch. & Mey. Ind. Sem. Hort. Petrop. 5: 36. 1838. 

 Heuchera hispida Hook. & Am. Bot. Beechey 347. 1838. Not Pursh, 1814. 

 Heuchera hirtifiora Torr. & Gray, Fl. N. Amer. 1 : 582. 1840. 

 Heuchera hemisphaerica Rydb. N. Amer. Fl. 22: 112. 1905. 

 Heuchera pilosella Rydb. loc. cit. 



Acaulescent or sometimes short-stemmed, flowering branches often more than one, scapi- 

 form or leafy, relatively thick, 2-6 dm. high, densely villous-hirsute with brownish gland- 

 tipped hairs. Basal leaves thick and often subcoriaceous, 4-7 cm. wide, cordate to rounded- 

 cordate, with mostly rounded lobes and somewhat doubly dentate with ovate apiculate teeth, 

 obtuse or acutish at the apex, hirsute on both sides, the hairs on the margin gland-tipped; 

 petioles 8-10 cm. long, densely covered with long glandular hairs ; cauline leaves similar though 

 smaller and shorter-petioled, often much reduced ; inflorescence relatively narrow, with the 

 exception of the flowers densely glandular, the cymules often very much contracted ; hypan- 

 thium hemispheric or at least rounded at the base, yellowish, often pink-tinged, densely pilose, 

 including the rounded or acutish sepals 3-4 mm. long; petals oblong-elliptic with a narrow 

 claw, longer than the sepals ; stamens exserted. 



Ocean bluffs and terraces, Humid Transition Zone; Humboldt County to San Luis Obispo County, California. 

 Type locality: probably Bodega Bay, Sonoma County, California. April-July. 



Heuchera Merriamii Eastw. Bull. Torrey Club 32: 199. 1905. Flowering branches several, short, 2 dm. 

 high or less, often bearing a small leaf at base. Basal leaves orbicular to ovate, truncate at base, 2-3 cm. wide, 

 their petioles 1-4 cm. long, glandular-hirsute; panicle contracted, glandular-puberulent; hypanthium hemispheric, 

 together with the sepals 3-4 mm. long. Apparently confined to the Salmon Mountain region of Trinity and Sis- 

 kiyou Counties, California. Type locality: Canyon Creek, Trinity County, California. 



2. Heuchera glabra Willd. Alpine Heuchera. Fig. 2291. 



Heuchera glabra Willd. ex Roem. & Sch. Syst. Veg. 6: 216. 1820. 

 Tiarella colorans Graham, Edinb. New Phil. Journ. 7: 349. 1829. 

 Heuchera divaricata Fisch. ex Ser. in DC. Prod. 4: 51. 1830. 



Flowering branches slender, erect or curved, 1-6 dm. high, glabrous, occasionally leafless, 

 more usually bearing 1-3 petiolate leaves, the smaller upper ones often subtending the lower 



