384 HYDRANGEACEAE 



those of the scape; hypanthium minutely glandular-puberulent, broadly turbinate, yellow with a 

 broad band of light brown above the middle, including the broadly triangular acute to obtuse 

 sepals 2.5-3 mm. long, in fruit becoming rounded at the base and greenish; petals little exceed- 

 ing the sepals, cream-colored, elliptic-oblong, mostly acuminate, the claw relatively broad ; sta- 

 mens not exceeding the sepals; styles short, ultimately exceeding the sepals by 0.5-1 mm. 



Rocky situations, Boreal Zones; White Mountains of eastern California. Type locality: McAfee Meadow, 

 White Mountains, Mono County, California. 



Family 60. HYDRANGEACEAE. 



Hydrangea Family. 



Trees, shrubs, or vines with simple opposite exstipulate leaves. Flowers corym- 

 bose, racemose or rarely solitary, perfect or the outer ones of the cluster sometimes 

 sterile. Hypanthium adnate to the ovary. Sepals and petals usually 5, sometimes 

 fewer. Stamens twice as many as the petals or more numerous. Ovary compound, 

 at least the lower half adnate to the hypanthium ; carpels 2-10, wholly united or 

 free at the apex. Seeds numerous ; embryo small ; endosperm copious. 



About 16 genera and 80 species, natives of temperate and tropical regions. 

 Capsule beaked by the persistent styles; ovules several to many in each cell, only 1 maturing in Fendlerella. 

 Styles more or less united; flowers showy. 



Leaves deciduous, often toothed; styles partly united. 1. Philadelphus. 



Leaves evergreen, coriaceous, entire; styles completely united. 2. Carpentaria. 



Styles distinct; capsule conical. 



Leaves toothed, small; stigma terminal; flowers showy. 3. Jamesia. 



Leaves entire, small; stigma partially introrse; flowers small. 4. Fendlerella. 



Capsule depressed-globose, beakless; styles distinct, deciduous; ovules solitary in each cell; flowers minute. 



5. Whipplea. 



1. PHILADELPHUS L. Sp. PI. 470. 1753. 



Branching shrub with deciduous, entire or toothed, usually petioled leaves. Flowers 

 perfect, usually showy and often fragrant, borne solitary or in few-flowered cymes at 

 the ends of short branchlets, often so grouped as to appear racemose or paniculate. 

 Sepals 4 or rarely 5, valvate, persistent. Petals 4 or 5, showy, white or ochroleucous, 

 convolute. Stamens usually numerous ; filaments subulate, free or united below ; anthers 

 short. Ovary adnate to the hypanthium for at least two-thirds its length, 4-celled, or 

 rarely 3- or 5-celled; styles elongated or short, more or less united. Ovules numerous, 

 pendulous in several series. Capsule obovoid, firm, loculicidal ; seeds numerous, with a 

 membranous testa; endosperm none. [Named after King Ptolemy Philadelphus.] 



A genus of about 45 species, natives of north temperate regions. Type species, Philadelphus coronarius L. 



Leaves 2.5-8 cm. long; flower clusters racemose or paniculate, 4-20-flowered ; calyx glabrous or nearly so without. 

 Leaves pubescent all over beneath, usually dentate; styles united for two-thirds their length. 



1. P. Gordonianus. 

 Leaves pubescent only on the nerves beneath, entire or denticulate. 



Styles united to the middle or only a little beyond. 2. P. Lewisii. 



Styles united up to the stigmas. 3. P. calif ornicus. 



Leaves 0.5-2 cm. long; flower clusters cymose, mostly 1-3-flowered; calyx silky-tomentose without. 



4. P. serpyllifolius. 



1. Philadelphus Gordonianus Lindl. Gordon's Syringa. Fig. 2303. 



Philadelphus Gordonianus Lindl. Bot. Reg. 24: Misc. 21. 1838. 

 Philadelphus columbianus Koehne, Gartenfl. 1896: 542. 1896. 

 Philadelphus angustifolius Rydb. N. Amer. Fl. 22: 166. 1905. 

 Philadelphus platyphyllus Rydb. N. Amer. Fl. 22: 167. 1905. 



Deciduous shrub, 2-4 m. high, with ascending branches. Leaves ovate, 3-8 cm. long, short- 

 acuminate, obtuse or on vigorous shoots subcordate at base, usually dentate, rarely sparsely 

 denticulate, more or less pubescent on both surfaces ; racemose flower clusters 5-10-flowered ; 

 petals oblong to oval, 15-20 mm. long; styles united half to two-thirds their length; capsule 

 ovoid, 8-10 mm. long. 



Hillsides and stream banks, mainly Humid Transition Zone; British Columbia, southward west of the 

 Cascades to Shasta and Humboldt Counties, California. Type locality: Northwest America, collected by Douglas. 

 May-July. 



2. Philadelphus Lewisii Pursh. Lewis' Syringa. Fig. 2304. 



Philadelphus Lewisii Pursh, Fl. Amer. Sept. 329. 1814. 

 Philadelphus confusus Piper, Bull. Torrey Club 29: 225. 1902. 

 Philadelphus Helleri Rydb. N. Amer. Fl. 22: 166. 1905. 



Deciduous shrub, 1-2.5 m. high, with ascending branches, the bark of the previous season 

 usually with numerous cross cracks and eventually exfoliating. Leaves ovate or ovate-lanceolate, 

 2-5 cm. long, acute at apex, obtuse or acute at base, entire or denticulate, glabrous or sparingly 



