EVENING-PRIMROSE FAMILY 185 



beakless to long-beaked. Seeds brown, sometimes somewhat cellular-puberulent, with fim- 

 briate upper margin. [Named for C. H. Godet, 1797-1879, author of Flora de Jura.] 



A genus of about 14 species, found in western North America and Chile. Most abundant in California. 

 Type species, Oenothera purpurea Curtis. 



Hypanthium with inner ring of hairs from one-fourth to three-fifths the way from the base to the summit; buds 

 erect, except in Numbers 3, 7, 8. 



Stigma-lobes less than 4 mm. long. 



Capsule 8-ribbed when immature, terete or somewhat square and 8-nerved when dried. 



Buds erect; sepals usually distinct, but sometimes united. 



Hypanthium 2-7 mm. long, tapered uniformly from base to summit; no annular swelling at 

 top of ovary. 



Inflorescence not congested in normal plants; capsules not enlarged at center; leaves 2-8 

 mm. wide. 1. G. quadrivulnera. 



Inflorescence congested in normal plants; capsule enlarged at middle; leaves 3-18 mm. 

 wide. 2. G. purpurea. 



Hypanthium S-IS mm. long, slender toward base and flaring out at summit; ovary usually 

 with annular swelling at top. 



Plants branching mainly from middle, erect; branchlets stout; capsules straight or nearly 

 so. 5. G. viminea. 



Plants branching from base, somewhat ascending; branchlets filiform; capsule usually 

 strongly arcuate. 6. G. parviftora. 



Buds nodding; sepals united in anthesis. 



Capsule rounded at base, not linear, with evident ribs in dry material; petals lavender without 

 purple base; stigma-lobes linear; inflorescence frequently glandular-pubescent. 



7. G. hispidula. 



Capsule elongate, linear, frequently square and smooth in mature specimens: petals lavender, 

 usually with purple base; stigma-lobes oval; inflorescence never glandular-pubescent. 



8. G. cylindrica. 



Capsule 4-sulcate when immature, terete and 8-nerved when mature. 3. G. amoena. 



Stigma-lobes 4-7 mm. long. 



Capsule not greatly enlarged at middle; petals not over 4 cm. long. 3. G. amoena. 



Capsule greatly enlarged at middle; petals 4-6 cm. long. 4. G. Whitneyi. 



Hypanthium with inner ring of hairs at least three-fifths of way from base to summit; buds nodding. 



Petals less than 1.5 cm. long, white or cream, or tinged with pink; stigma-lobes very short, less_ than 0.5 



mm. long. 1 1 . C epilobioides. 



Petals 1.5 cm. or more long, lavender to purple; stigma-lobes more than O.S mm. long. 



Capsule very narrowly linear, 2-4 cm. long, 1-2 mm. thick; petals lavender with purple base. 



8. G. cylindrica. 

 Capsule thicker; petals lavender, but without purple base. 



Filaments unequal, slightly flattened; living capsule terete and faintly nerved, mature and dry 

 capsule square and obscurely nerved or smooth; pedicels as much as 2 cm. long 



9. G. Bottae. 



Filaments subequal, subfiliform; capsule 8-ribbed, these ribs especially evident in dry or mature 

 material ; pedicel usually less than 3 mm. long. 

 Petals not bilobed; capsule sessile or nearly so. 10. G. Dudteyana. 



Petals bilobed; capsule with short pedicel. 12. G. biloba. 



1. Godetia quadrivulnera (Dougl.) Spach. Four-spotted Godetia. Fig. 3396. 



Oenothera quadrivulnera Dougl. ex Lindl. Bot. Reg. 13: pi. 1119. 1828. 

 Godetia quadrivulnera Spach, Hist. Veg. 4: 389. 1835. 

 Godetia hingensis Suksd. Deutsch. Bot. Monatss. 18:88. 1900. 

 Godetia Goddardii Jepson, Univ. Calif. Pub. Bot. 2: 341. 1907. 

 Godetia sparsifolia Jepson, loc. cit. 



Usually erect, branching from base or near middle, 1-8 dm. tall. Leaf-blades lanceolate to 

 oblong, 2-5 cm. long, acute, sessile or nearly so; buds erect; hypanthium 2-6 mm. long, with 

 inner ring of hairs about one-third way from base ; sepals green or yellow, usually distinct in 

 anthesis, 5-10 mm. long; petals lavender to purple, with or without darker spot near center, 

 5-20 mm. long, cuneate; capsules 1-3.5 cm. long, 2-3 mm. thick, terete and 8-ribbed with a faint 

 nerve between the ribs when fresh, or square and conspicuously ribbed when dry, the ribs all 

 about equally prominent, sessile or on pedicels as much as 2 mm. long, tapering into a beak 

 . 5-2 mm. long ; seeds 1 mm. long, equally broad, with cresting about one-fifth as long as body 

 of seed. 



Open hillsides, particularly at edge of woods, Upper Sonoran and Transition Zones; Washington to Lower 

 California. Type locality: "north-west of North America." Collected by Douglas. April-July. 



Godetia quadrivulnera var. Davyi Jepson, Univ. Calif. Pub. Bot. 2: 341 1907. Leaves short, 1-2 cm. 

 long, oblong to spatulate; capsules scarcely beaked, ribs very prominent, especially along the sutures. Wear tbe 

 coast from Del Norte County to Monterey County, California. Type locality: Point Reyes, Mann County. 



Godetia quadrivulnera var. vacensis Jepson, loc. cit. Leaves lanceolate, 2-5 cm long; sepals usually 

 united in anthc-sis; capsule 2-4 cm. long, very slender, with a beak 2-4 min. long, capsule-ribs of equal pronii- 

 nence. At scattered stations in California from Solano and Sacramento Counties to Ventura County, lype 

 locality: Vaca Mountains. 



2. Godetia purptirea (Curtis) G. Don. Purple Godetia. Fig. 3397. 



Oenothera purpurea Curtis, Bot. Mag. 10: pi. 352. 1796. 

 Godetia Willdenowiana Spach, Hist. Veg. 4: 387. 1835. 

 Godetia purpurea G. Don in Sweet, Hort. Brit. ed. 3. 237. 1839. 



Erect, simple or with branches closely crowded at summit, 1-6.5 dm. tall, glabrate to pubes- 

 cent. Leaves broadly elliptic, 2-4 cm. long, 1-2 cm. wide, puberulent yet appearing glaucous, tne 



