EVENING-PRIMROSE FAMILY 193 



1. Oenothera Rydbergii House. Rydberg's Evening-primrose. Fig. 3408. 



Onagra strigosa Rydb. Mem. N.Y. Bot. Gard. 1:278. 1900. 



Oenothera biennis var. strigosa Piper in Piper & Beattie, Fl. Palouse Reg. 124. 1901. 



Oenothera strigosa Mack. & Bush, FI. Jackson Co. Mo. 139. 1902. Not O. strigosa Willd., a herbarium name 



as synonym in Sprengel, Syst. 2: 228. 1825. 

 Oenothera Rydbergii House, N.Y. State Mus. Bull. No. 233-234: 61. 1921. 



Biennial, grayish-strigose throughout, erect, largely unbranched, 3-10 dm. tall, strigose and 

 hirsute, sometimes with reddish tinge. Lowest leaves spatulate, obtuse, 3-10 cm. long, 1-2 cm. 

 wide, with petioles an additional 1-3 cm. long, these leaves passing gradually into lanceolate, 

 acute, repand-denticulate leaves of the stem, with shorter petioles ; inflorescence with leafy lanceo- 

 late, subsessile bracts 1-5 cm. long ; flowers vespertine ; bracts longer than mature capsules ; hy- 

 panthium 3^ cm. long, pubescent within, often hirsute without; sepals strigose and hirsute, 10-15 

 mm. long, with free tips 2 mm. long; petals yellow, broadly obcordate, 1.2-2 cm. long; stamens 

 quite equal, about as long as petals, glabrous ; style pubescent on lower portion ; stigma-lobes 

 5-7 mm. long; capsule 2.5-3.5 cm. long, tapering slightly; seeds reddish brown, obtusely angled, 

 irregular, 1-1.5 mm. long. 



Moist places, meadows, stream banks. Upper Sonoran and Transition Zones; from eastern Washington and 

 Oregon to Minnesota and Kansas. Type locality: Pony, Montana. July-Aug. 



Oenothera cheradophila Bartlett, Bot. Gaz. 44: 302. 1907. Not certainly distinct from O. Rydbergii, 

 differing by: sepal-tips 1 mm. long; petals about 8 ram. long; floral bracts usually much shorter than mature 

 capsules. Southeastern Washington. Type locality: Bingen, Washington. 



2. Oenothera biennis L. Small-flowered Evening-primrose. Fig. 3409. 



Oenothera bicnttis L. Sp. PI. 346. 1753. 



Much like the preceding species, simple or branched, stems reddish, finely pubescent and hir- 

 sute, the larger hairs from reddish pustules. Cauline leaves broadly lanceolate to narrowly rhom- 

 boid ; flowers and buds in dense corymbose clusters ; sepals with free tips 3-4 mm. long ; petals 

 12-15 mm. long. 



Moist places, Transition Zones; western Washington and Oregon into Canada, then east to Atlantic Coast. 

 Type locality: "Virginia." July-Aug. 



3. Oenothera Hookeri Torr. & Gray. Hooker's Evening-primrose. Fig. 3410. 



Oenothera Hookeri Torr. & Gray, Fl. N. Amer. 1 : 493. 1840. 

 Oenothera Jepsonii Greene, Fl. Fran. 211. 1891. 

 Onagra Hookeri Small, Bull. Torrey Club 23: 171. 1896. 

 Oenothera franciscana Bartlett, Rhodora 15: 35. 1914. 



Perennial, or possibly short-lived perennial, largely with spreading pubescence ; the stems 

 hirsute, muricate with reddish pustules, simple and erect, or somewhat branched mostly from the 

 base, 6-12 dm. tall. Lower leaves oblanceolate to spatulate with blades 5-20 cm. long, on petioles 

 half as long ; cauline leaves narrowly to broadly lanceolate, on shorter petioles, sinuate-denticu- 

 late, green, wavy, soft-hairy, gradually reduced up the stem to the leafy bracts of the elongate 

 inflorescence; hypanthium 3.5-4 cm. long, pubescent within; sepals red, densely hirsute, conspicu- 

 ously papillose at base of longer hairs, 22-25 mm. long ; petals 25-35 mm. long, yellow but turn- 

 ing reddish in age, broadly obovate; stamens equal, two-thirds as long as petals; stigma-lobes 

 4-6 mm. long ; capsule obtusely quadrangular, 2-5 cm. long, hirsute ; seeds reddish brown, sharply 

 angled, 1.5 mm. long. 



Moist places, Upper Sonoran and lower Transition Zones; largely from Lake and Sutter Counties to San 

 Luis Obispo County, California. Type locality: California, probably San Francisco. June-Sept. 



Oenothera Hookeri subsp. montereyensis Munz, El Aliso 2: 14. 1949. Plants bushy; flowers large; buds 

 blunt, the sepal-tips 1-2.5 mm. long; sepals with short gland-tipped hairs and papillose at base of longer hairs. 

 Sea cliffs, central California, San Mateo County to San Luis Obispo County. Type locality: Alder Creek, 

 Monterey County. 



Oenothera Hookeri subsp. Wolfii Munz, op. cit. 16. Flowers small, the petals 2-2.5 cm. long; sepals 

 glandular-pubescent and papillose at base of longer hairs; sepal-tips 2-3 mm. long. In sand or gravel along 

 streams, Jackson County, Oregon, to Trinity and Humboldt Counties, California. Type locality: near Trinidad, 

 Humboldt County. 



Oenothera Hookeri subsp. veniJsta Munz, op. cit. 21. {Oenothera vcnusta Bartlett, Rhodora 16: 36. 

 1914.) Whole plant rather grayish; stems 1.5-2 m. tlal, freely branched throughout, hirsute and' muricate; 

 leaves soft-hairy, wavy; sepals green, sparsely hirsute, scarcely papillose at base of hairs; petals 3-4 cm. long; 

 capsule hirsute. Moist places. Upper Sonoran and lower Transition Zones below 4,500 feet, central interior to 

 southern California. Type locality: San Bernardino, California. 



Oenothera Hookeri subsp. ornata (A. Nels.) Munz, op. cit. 25. {Onagra ornata A. Nels. Bot. Mag. 

 52: 268. 1911.) Stem-leaves plane; sepals pilose and glandular-pubescent; sepal-tips 2.5-4 mm. long; seeds 

 about 1.3 mm. long. Eastern Washington and western Idaho. Type locality: Boise, Idaho. 



Oenothera Hookeri subsp. angustifolia (Gates) Munz, op. cit. 26. {Oenothera Hookeri var. angustifolia 

 Gates, Mut. Factor in Evol. 10, 30. 1915.) Stems reddish, 3-9 dm. high, somewhat muncate, simple or tew- 

 branched mostly from base; leaves plane, green, not conspicuously soft-hairy; sepals red, ,with long spreading 

 hairs and short gland-tipped ones, scarcely or not at all papillose. Transition Zone, mostly above 5,10U teet, 

 eastern Washington to southern California and east to Idaho, Colorado, and New Mexico. Type locality: 

 Asphalt, LUah. 



Oenothera Hookeri subsp. grisea Munz, op. cit. 29. {Oenothera venusta var. grisea Bartlett, Rhodora 



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