194 ONAGRACEAE 



X Oenothera erythrosepala Borb. Magyar Bot. Lapok. 2: 245. 1903. Very near to O. Hookeri and 

 differing from it by the broader, more crinkled leaves, the upper ones sessile, by the broad floral bracts, and 

 by the perhaps paler yellow petals, there is an escape along the northern California coast to western Washington. 

 It is the plant distributed by De Vries as O. Latnarckiana, but is not O. Lamarkiana Ser. 



Oenothera longissima subsp. Glutei (A. Nels.) Munz, op. cit. 46. {Oenothera Glutei A. Nels. Amer. 

 Bot. 28: 22. 1922.) Like O. Hookeri, but with hypanthium 8-12 cm. long. California (eastern Mojave Desert) 

 to southern Utah. Type locality: Navajo Mountains, Coconino County, Arizona. 



Oenothera stricta Ledeb. in Link, Enum. Hort. Berol. 1: 377. 1821. (Oenothera arguta Greene, Fl. Fran. 

 212. 1891.) Decumbent perennial about 3 dm. tall with linear-lanceolate, saliently dentate leaves, the cauline 

 broadest at the sessile, somewhat clasping base. A South American species growing spontaneously on the south- 

 ern shore of Monterey Bay, California. 



Oenothera laciniata Hill, Hort. Kew. 172. pi. 6. 1769. This species of the subgenus Raimannia can be 

 distinguished by its erect buds; petals pale yellow, about 1 cm. long; seeds not angled, yellowish, pitted; capsule 

 narrowly cylindrical; leaves sinuate-dentate or pinnatifid. Native of the southern and eastern states and spar- 

 ingly naturalized, as at Pasadena and Banning, California. Type locality: Carolina. 



Oenothera speciosa Nutt. Journ. Acad. Phila. 2: 119. 1821. This species is of the subgenus Hartmannia 

 and is characterized by having leaves lanceolate, sinuate or pinnatifid; flowers large, white, the petals 3-4 cm. 

 long; capsules 4-winged and 4-ribbed, clavate, 1-1.5 cm. long. Native from Arizona eastward. Sparingly nat- 

 uralized as at Pomona, California. A pink-flowered form (Oenothera speciosa var. Chitdsii (Bailey) Munz, 

 Leaflets W. Bot. 2: 87. 1938.) native along the gulf coast of Texas is in common cultivation as Mexican 

 Evening-primrose. 



4. Oenothera deltoides Terr. & Frem. Large Desert Evening-primrose. 



Fig. 3411. 



Oenothera deltoides Torr. & Frem. in Frem. Second Rep. 315. 1845. 

 Oenothera trichocalyx of authors for much of our material, not Nutt. 



Coarse spring or winter annuals, simple or more frequently with central erect stem, 5-25 cm. 

 tall, and few to several decumbent branches naked at the base and 5-100 cm. long; stems pale 

 green, with exfoliating epidermis, glabrous in lower parts, with spreading hairs in upper parts. 

 Lower leaves in sort of rosette, the blades rhombic-obovate to rhombic-lanceolate or oblanceo- 

 late, subentire to remotely denticulate or even dentate, 2-8 cm. long, narrowed into slightly 

 winged petioles of same or less length ; cauline leaves gradually somewhat reduced, becoming ses- 

 sile and dentate, but not pinnatifid; flowers solitary in axils, vespertine, buds nodding, obtuse; 

 hypanthium slender, 2-4 cm. long, it and buds having straight spreading hairs 1-1.5 mm. long; 

 sepals lance-linear, 15-35 mm. long; petals white, turning pink with age, 2-4 cm. long; 

 stamens subequal ; stigma-lobes Z-6 mm. long; capsules spreading, even reflexed, woody, with 

 exfoliating epidermis, prismatic-cylindric, 4-5 (7) cm. long, commonly 2-3 mm. thick at base ; 

 seeds narrowly obovoid, 1.5-2 mm. long, light brown, usually with purple spots and rows of 

 minute cellular pitting. 



Sandy places. Lower Sonoran Zone; deserts of southern California and adjacent Arizona. Type locality not 

 given. March-May. 



Oenothera deltoides var. HowelHi Munz, El AHso 2: 81. 1949. Leaves runcinate-pinnatifid, lanceolate, 

 3-12 cm. long, 1-3 cm. wide, cinereous; buds acute. Sand dunes, Antioch, Contra Costa County, California. 



Oenothera deltoides var. Piperi Munz, Amer. Journ. Bot. 18: 314. 1931. Plants low, frequently less 

 than 1 dm. tall and often simple; upper leaves lanceolate in outline and deeply and regularly sinuate-dentate to 

 pinnatifid, with rachis 3-4 mm. wide, well provided with long soft curly hairs, about 2 mm. long; ovary and 

 sepals with same hairs; petals usually less than 2 cm. long; capsules 1.5-3 cm. long, 3-5 mm. thick at base. 

 Upper Sonoran Zone from eastern Oregon and northeastern California and western Nevada. Type locality: 

 Man's Lake, eastern Oregon. 



Oenothera deltoides var. cognata (Jepson) Munz, op. cit. 313. Plants commonly 2-4 dm. tall and branched 

 from base; upper leaves coarsely sinuate-dentate but rarely pinnatifid, blades 5-10 mm. wide; hair on upper parts 

 as in Piperi; petals 2.5-3.5 cm. long; capsules 2.5-7 cm. long, 3-5 mm. thick at base. Sandy plains. Upper 

 Sonoran Zone; Great Valley of California, and western end of Mojave Desert. Type locality: Corral Hollow, 

 Alameda County, California. 



Oenothera deltoides var. cinerJcea (Jepson) Munz, op. cit. 316. Habit, foliage, and flowers as in the 

 species, but hair short, less than 1 mm. long and closely appressed. Sandy desert. Lower Sonoran Zone, southern 

 part of California and adjacent Arizona. Type locality: Borrego Springs, San Diego County, California. 



5. Oenothera pallida Lindl. Pallid Evening-primrose. Fig. 3412. 



Oenothera pallida Lindl. Bot. Reg. 14:/)/. 1142. 1828. 

 Anogra Douglasiana Spach, Ann. Mus. Paris IL 4: 339. 1835. 

 Oenothera leptophylla Nutt. ex S. Wats. Proc. Amer. Acad. 8: 602. 1873. 

 Anogra pallida Britt. Mem. Torrey Club 5: 234. 1894, in part. 



Strongly rooted perennial, with creeping rootstalks, with main stem erect, 2-5 dm. tall, and 

 several spreading or ascending branches, epidermis white, quite glabrous and exfoliating, or with 

 few scattering long hairs in upper parts. Cauline leaves mostly lanceolate to lance-linear, suben- 

 tire to remotely denticulate or sinuate-dentate, usually with undulate margin ; the blades 2-6 cm. 

 long, sessile or short petioled ; flowers vespertine, fragrant ; buds acuminate, nodding ; hypan- 

 thium very slender, frequently reddish, 2-3.5 cm. long, usually glabrous; sepals 12-18 mm. long, 

 the free tips 0.5-2 mm. long; petals white, turning pink, broadly obovate, 1-3 cm. long; stamens 

 subequal, glabrous ; capsule usually curved, often somewhat contorted, glabrate, 1 . 5-4 cm. long, 

 subcylindric, 2-3 mm. thick at base, tapering gradually toward apex ; seeds in 1 row, linear-obo- 

 void, 1 . 5-2 mm. long, brown with dark spots or quite dark, minutely pitted under strong lens. 



Sandy places and dry plains. Upper Sonoran Zone; eastern Washington and Oregon to Utah and New 

 Mexico. Type locality: "north-west of North America." Collected by Douglas. May-Aug. 



