208 ONAGRACEAE 



the alternate set much reduced and usually sterile. Stigma capitate. Capsule 2-celled, 4- 

 valved, linear or clavate. Seeds many, in a single row in each cell, not comose. [Name 

 in honor of Gay, author of Flora of Chile, and Greek word for plant.] 



A genus of 9 species of the temperate regions of western North and South America. Type species, 

 Gayophytum humile Juss. 



Capsule torulose, pedicelled; plants freely branched above the base, repeatedly dichotomous, the upper leaves 

 bract-like. 

 Seeds glabrous. 



Petals O.S-1.5 mm. long. 



Petals 0.5 mm. long; capsule 2-5 mm. long, shorter than the deflexed pedicel; plants quite glabrous. 



1. G. ramosisstmum. 



Petals 1-1.5 mm. long; capsule S-12 mm. long, exceeding the pedicel. 2. G. Nuttallii. 



Petals 2-4 mm. long. 3. G. diffusum. 

 Seeds appressed-canescent. 



Petals 1-2 mm. long. 4. G. lasiospermum. 



Petals 3-4 mm. long. 5. G. eriospermum. 



Capsule not torulose, subsessile; plants branched mostly at the base, not so much above; upper leaves quite well 

 developed. 

 Seeds vertically placed in a very narrow capsule. 



Seeds glabrous. 6. G. racemosum. 



Seeds appressed-canescent. 7. G. Helleri. 



Seeds obliquely placed in a slightly broader capsule. 8. G. humile. 



1. Gayophytum ramosissimum Torr. & Gray. Much-branched Gayophytum. 



Fig. 3445. 



Gayophytum ramosissimum Torr. & Gray, Fl. N. Amer. 1: 513. 1840. 

 Gayophytum ramosissimum var. deflexum Hook. Lond. Journ. Bot. 6: 224. 1847. 

 Gayophytum ramosissimum var. obtusum Jepson, Man. Fl. PI. Calif. 688. 1925. 



Diffusely branched, mostly above the base, with the ultimate branches filiform, quite glabrous, 

 sometimes slightly strigillose about the flowers ; plant 2-5 dm. tall. Leaves lance-linear, 1-3 cm. 

 long, short-petioled, gradually reduced up the stem ; pedicels capillary, 3-5 mm. long, mostly 

 spreading-deflexed ; flowers minute; sepals erect, 0.5 mm. long; petals 0.5 mm. long; stigma 

 globose; capsule plump, 2-5 mm. long; seeds glabrous, 0.6 mm. long. 



Dry slopes and ridges. Upper Sonoran and Transition Zones; from eastern Washington and Oregon to 

 northeastern California, Wyommg, Utah, and northern Arizona. Type locality: "Rocky Mountains," Blackfoot 

 River. June-Aug. 



2. Gayophytum Nuttallii Torr. & Gray. Nuttall's Gayophytum. Fig. 3446. 



Gayophytum Nuttallii Torr. & Gray, Fl. N. Amer. 1: 514. 1840. 



Gayophytum ramosissimum var. strictipes Hook. Lond. Journ. Bot. 6:224. 1847. 



Gayophytum ramosissimum var. pygmaeum Jepson, Man. Fl. PI. Calif. 689. 1925. 



With habit and stature of G. ramosissimum, usually more evidently strigillose in the upper 

 parts; pedicels 1-3, rarely 5 mm. long, erect; sepals 1-1.5 mm. long, reflexed in anthesis ; petals 

 reddish, at least in age, 1-1.5 mm. long; capsules 5-12 mm. long, erect, usually exceeding the 

 pedicels; seeds glabrous, 1-1.5 mm. long. 



Dry slopes and ridges, Upper Sonoran and Transition Zones; Washington to southern Cailifornia, Dakota, 

 and New Mexico; Chile and Argentina. Type locality: "Rocky Mountains." June— Aug. 



Gayophytum Nuttallii var. intermedium (Rydb.) Munz, Amer. Journ. Bot. 19: 772. 1932. (Gayophy- 

 tum intermedium Rydb. Bull. Torrey Club 31: 569. 1904.) Puberulence appressed; pedicels and fruit 

 spreading or deflexed. Transition Zone, eastern Washington and Oregon to Wyoming and Colorado. Type 

 locality: Ouray, Colorado. 



Gayophytum Nuttallii var. Abramsii Munz, loc. cit. Puberulence short and spreading; pedicels and cap- 

 sules mostly erect. Dry slopes. Transition Zone; eastern Washington and Oregon to southern California, Mon- 

 tana, and Nevada. Type locality: Coldwater Canyon, San Antonio Mountains, Los Angeles County, California. 



3. Gayophytum diffiisum Torr. & Gray. Diffuse Gayophytum. Fig. 3447. 



Gayophytum diffusum Torr. & Gray, Fl. N. Amer. 1: 513. 1840. 



With general habit and stature of G. Nuttallii; appressed-puberulent in upper parts ; pedicels 

 2--8 mm. long, erect or divaricate ; sepals 2-3 mm. long, reflexed in anthesis ; petals white to 

 pink, 2-4 mm. long ; stigma clavate-capitate ; capsules 5-12 mm. long, divaricate ; seeds glabrous, 

 1—1 .25 mm. long. 



Occasional on dry slopes. Transition Zone; Washington to southern California, Montana, and Wyoming. 

 Type locality: "Rocky Mountains and plains of Oregon." June-July. 



Gayophytum diffusum var. villosum Munz, Amer. Journ. Bot. 19: 773. 1932. Upper parts of plant with 

 a short spreading pubescence. Rare, Washington to mountains of southern California and Idaho. Type locality: 

 Farewell Bend, Crook County, Oregon. 



4. Gayophytum lasiospermum Greene. Hairy-seeded Gayophytum. Fig. 3448. 



Gayophytum lasiospermum Greene, Pittonia 2: 164. 1891. 



Plant 2-5 dm. high, branching freely above, with appressed puberulence in the upper parts. 

 Leaves linear to lance-linear, 1-3 cm. long, entire ; pedicels 3-6 mm. long, divaricate to spreading 



