430 POLEMONIACEAE 



times the throat and tube dark; stamens inserted about midway on throat, anthers barely ex- 

 serted from orifice of throat, filaments glabrous ; stigmas included or shortly exserted ; styles 

 included (except in subspecies) ; capsule ellipsoid, locules several-seeded, seeds beset with rugose 

 papillae. A highly polymorphic genetic complex. 



Grassy hills and open woods. North Coast Ranges of California from Monterey County to Humboldt County. 

 Type locality: "California." Collected by Douglas. April-June. 



Key to Subspecies. 

 Style and usually also the stigma included. 



Papillae of seed-coat with no definite pattern, rarely indistinctly annular; stigma-lobes usually included, rarely 

 exserted. 

 Corolla-limb 10-15 mm. broad, tube 1-2 mm. broad. 

 Corolla-tube 1 .5-2 times the calyx. 



Stems simple or with few erect branches from base; Monterey County to Humboldt County in 



open woods and grassland, typical. L. androsaceus. 



Stems much-branched, spreading or decumbent, sometimes erect; Monterey County south in 

 Coast Ranges to the Tehachapi and San Gabriel Mountains. subsp. Plaskettii. 



Corolla-tube 3-4 or 5 times the calyx, limb sometimes as much as 15 mm. broad; plants spreading or 

 decumbent; coastal bluffs and mesas from Sonoma County south to Monterey County. 



subsp. croceus. 



Coralla-limb 6-10 mm. broad, the tube usually filiform or at least slender; Inner and Outer Coast 

 Ranges; Lake and Mendocino Counties to Santa Barbara County. subsp. hUeus. 



Papillae of seed-coat in a definite corrugated annular pattern around the seed; stigma-lobes exserted or in- 

 cluded. 

 Corolla always white; stems simple and erect or with 2 or 3 erect stems; Sierra Nevada foothills from 



Fresno County to Butte County. subsp. laetus. 



Corolla usually yellow, pink, or lilac, less commonly white, stems decumbent or ascending; southern 

 California north in the Inner South Coast Ranges and the hills bordering the southern Salinas 

 Valley to southern Monterey County. subsp. Ititeotus. 



Style and stigma long-exserted; papillae of seed-coat forming an annular pattern; plants usually erect, simple or 

 with several erect stems; usually on the coastal slopes of the mountains of southern California north along 

 coast to Monterey Bay. subsp. micranthus. 



Linanthus androsaceus subsp. Plaskettii (Eastw.) H. L. Mason. (Linanthus Plaskettii Eastw. Bot. Gaz. 

 37: 443. 1904; L. graciosus Milliken, Univ. Calif. Pub. Bot. 2: 59. 1904; Cilia graciosa Brand, Pflanzenreich 

 4250. 140. 1907; G. tassajarae Brand (orth. emend.) loc. cit., as G. jassajarae.) Similar to the typical species 

 except for its slightly smaller corollas, much-branched and spreading habit, and more southern range. Type 

 locality: "Santa Lucia Mts." 



Linanthus androsaceus subsp. croceus (Milliken) H. L. Mason. (Gilia longituba Benth. PI. Hartw. 324. 

 1849; Leptosiphon parviflorus var. rosaceus Hook. Bot. Mag. 96: pi. 5863. 1870; Gilia lutea var. rosea Regel, 

 Gartenfl. 20: 97. pi. 682. 1871; Leptosiphon roseus Thompson, Gard. Chron. 1264. 1871; Gilia androsacea var. 

 rosacea A. Gray, Bot. Calif. 1: 491. 1880; Linanthus rosaceus Greene, Pittonia 2: 259. 1892; L. parviflorus var. 

 rosaceus Jepson, Fl. \V. Mid. Calif. 431. 1901 ; L. parviflorus var. croceus Milliken, Univ. Calif. Pub. Bot. 2: 59. 

 May, 1904; L. croceus Eastw. Bot. Gaz. 37: 449. June, 1904; L. longituba Heller, Muhlenbergia 1: 43. 1904; 

 Cilia longituba subsp. eu-longituba Brand, Pflanzenreich 4^": 140. 1907; G. longituba var. rosacea Brand, loc. 

 cit.; G. longittiba subsp. crocca Brand, op. cit. 141.) Type locality: "Monterey." 



Linanthus androsaceus subsp. luteus (Benth.) H. L. Mason. {Leptosiphon luteus Benth. Bot. Reg. 19: 

 under pi. 1622. 1833; Cilia lutea Steudel, Nom. ed. 2. 1: 684. 1840; G. micrantha var. atirea Benth. PI. Hartw. 

 325. 1849; G. lutea subsp. eu-lutea Brand, Pflanzenreich 4^": 142. 1907; C. lutea var. aurea Brand, op. cit. 143; 

 Linanthus parviflorus of Jepson, Man. FI. PI. Calif. 805, in part. 1925; Fl. Calif. 213. 1943.) Type locality: 

 "California." Collected by Douglas. 



Linanthus androsaceus subsp. laetus H. L. Mason, Madrono 9:249. 1948. Type locality: Cherokee, 

 Butte County. 



Linanthus androsaceus subsp. luteolus (Greene) H. L. Mason. Linanthus luteotus Greene, Erythea 3: 121. 

 1895; L. parviflorus var. luteolus Milliken, Univ. Calif. Pub. Bot. 2: 58. 1904; Cilia lutea subsp. luteola Brand, 

 Pflanzenreich 4^"; 143 1907.) Type locality: Cuyamaca Mountains, San Diego County. 



Linanthus androsaceus subsp. micranthus (Steudel) H. L. Mason. {Leptosiphon parviflorus Benth. Bot. 

 Reg. 19: under pi. 1622. 1833; Gilia micrantha Steudel ex Benth. A. DC. Prod. 9: 315. 1845; Linanthus parvi- 

 florus Greene, Pittonia 2: 258. 1892; Gilia lutea subsp. micrantha Brand, Pflanzenreich 4260; 142. 1907; G. lutea 

 var. longistylus Munz, Man. S. Calif. 398. 1935.) Type locality: "California." Collected by Douglas. 



31. Linanthus bicolor (Nutt.) Greene. Bicolored Linanthus. Fig. 3956. 



Leptosiphon bicolor Nutt. Journ. Acad. Phila. II. 1: 156. 1848. 



Gilia tenella Benth. PI. Hartw. 325. 1849. Not G. tenella A. Gray or Nutt. 



Navarretia tenella Kuntze, Rev. Gen. PI. 2: 433. 1891. 



Linanthus bicolor Greene, Pittonia 2: 260. 1892. 



Linanthus Eastwoodiae Heller, Muhlenbergia 1 : 125. 1905. 



Gilia bicolor Piper, Contr. U.S. Nat. Herb. 11: 460. 1906. 



Linanthus diffusa Heller ex Brand, Pflanzenreich 42=0: 139, as a synonym. 1907. 



Erect or spreading annual, 5-16 cm. high, simple or branched, herbage puberulent to hispid. 

 Leaves 3-7-cleft into linear segments or the middle segment broader, then from orbicular to 

 linear ; flowers sessile in densely bracteate heads, rarely more than one or two blooming at a 

 time in each head ; calyx deeply cleft to the very short tube, hispid, the sinuses usually with a 

 very small inconspicuous membrane, rarely naked; corolla salverform, 15-30 mm. long, always 

 bicolored, the lobes from red to pink or white, throat yellow, tube 2-A times the calyx, stout, 

 throat ample, 1-3 mm. long ; stamens inserted in the throat, exserted from the orifice ; stigma 

 0.5-1.5 mm. long; capsule-locules few-seeded. 



Vancouver Island, British Columbia, south through the Cascade Mountains of Oregon and Washington to the 

 Coast Ranges and Sierra Nevada foothills, California, coastal and insular southern California. Type locality: "on 

 the Oregon near the outlet of the Wahlamet." April-June. 



Linanthus bicolor subsp. minimus H. L. Mason, Madrono 9: 250. 1948. Corolla minute, about 1 cm. long, 

 white to sordid. Coastal area from Bodega Head, California, north to Puget Sound, Washington. Type locality: 

 Gages Point, Skagit County, Washington. 



