BORAGE FAMILY 555 



ing hairs as are the petioles. Lower leaves 10-20 cm. long, oblong-oblanceolate to narrowly 

 spatulate, acute or rounded at apex, blade gradually narrowed to a petiole of about equal or 

 shorter length, 1-2.5 cm. wide; upper stem-leaves sessile or subsessile, linear-oblong to broadly 

 lanceolate, acute or acutish, all rather sparsely strigose ; inflorescence at first narrow becoming 

 loose and widely branched in age; calyx-lobes ovate, 2-2.5 mm. long; corolla white, the tube 

 equaling the calyx-lobes, limb 6-10 mm. broad; appendages broader than long, puberulent; 

 nutlets 5 mm. long, the dorsal face dull, muriculate, covered with subequal barbed prickles. 



Open forests and edges of mountain meadows, Transition and Canadian Zones; southern Oregon in Jack- 

 son and Klamath Counties, south in the Coast Ranges to Humboldt and Lake Counties, and in the Sierra 

 Nevada to Alpine County, California. Type locality: Mount Shasta, California. June-Aug. 



15. Hackelia mundula (Jepson) Ferris. Pink Stickseed. Fig. 4216. 



Lappula calif ornica var. mundula Jepson, Fl. Calif. 3: 309. 1943. 



Stems 1 to several from a woody root, 5-9 dm. high, more or less velvety-pubescent through- 

 out. Leaves oblanceolate, acute or obtuse at apex, 8-15 cm. long, stem-leaves sessile, reduced 

 upward, the bracts tending to be broad and clasping; inflorescence with spreading branches, 

 widely divaricate in fruit, fruiting pedicels 8-14 mm. long; flowers pink with whitish centers, 

 fading blue, 12-20 mm. broad; tube about equaling the calyx, crests conspicuous, erect-spreading, 

 narrowly ovate, slightly emarginate, puberulent ; nutlets 6-8 mm. long, the dorsal face shining, 

 smooth, prickles slender, 2.5-5 mm. long. 



North slopes in red fir forests, Canadian Zone; in the Sierra Nevada from Yosemite National Park to 

 Tulare County, California. Type locality near Long Meadow, Tulare County. June-July. 



16. Hackelia velutina (Piper) I. M. Johnston. Velvety Stickseed. Fig. 4217. 



Lappula velutina Piper, Bull. Torrey Club 29: 546. 1902. 



Hackelia velutina I. M. Johnston, Contr. Gray Herb. No. 68: 47. 1923. 



Hackelia loiigituba I. M. Johnston, Journ. Arnold Arb. 29: 237. 1948. 



Stems 1 to several from the perennial root, 3-8 dm. high, clothed throughout with densely 

 velvety pubescence. Lower leaves spatulate-oblanceolate, acutish to obtuse, 8-14 cm. long, 

 petiole about as long or longer than the blades ; cauline leaves scattered or often numerous, 

 oblong-linear to lanceolate, sessile with broad, often suborbicular bases, acute or obtuse at apex, 

 gradually reduced upwards, 2-8 cm. long ; inflorescence rather compact, spreading in fruit, the 

 branches 5-10-flowered, fruiting pedicels 5-10 mm. long; calyx-lobes oblong, obtuse, densely 

 canescent; corolla blue, tube 5 mm. long, well-exceeding the calyx, limb 12-20 mm. broad; 

 appendages spreading, white, conspicuous, approximately one-third as long as the corolla-lobes, 

 the deeply 2-lobed crests pilose, equaled in length by the oblong protuberance; nutlets 6-8 mm. 

 long, the dorsal face dull, muriculate, beset with rather slender barbed prickles 2-3.5 mm. long 

 and rather thinly puberulent. 



Dry open forests, mainly Transition and Canadian Zones; Sierra Nevada, from Plumas County to Tulare 

 County, California. Type locality: "General Grant Grove," Tulare County. June-Aug. 



17. Hackelia nervosa (Greene) I. M. Johnston. Sierra Stickseed. Fig. 4218. 



Echinospcrmuni nervosum Kell. Proc. Calif. Acad. 2 : 146. fig. 42. 1862. 



Lappula nervosa Greene, Pittonia 2: 182. 1891. 



Hackelia nervosa \. M. Johnston, Contr. Gray Herb. No. 68: 47. 1923. 



Stems 1 to several, erect, 2-5 dm. high, glabrous or pilose with spreading hairs. Basal leaves 

 oblong-spatulate to linear-spatulate, narrowed to more or less winged petioles of about equal 

 length, lower stem-leaves shorter, subsessile and acutish, those subtending the lower branches 

 of the inflorescence ovate-lanceolate, subclasping at base, very acute or short-acuminate at apex, 

 retrorsely appressed-pubescent beneath, strigose above, often thinly so on the upper leaves, 

 harshly pubescent in age; racemes slender, 2-10-flowered ; pedicels slender, at length spreading 

 or recurved; calyx-lobes oblong to oblong-ovate, 2.5-3 mm. long; corolla blue, the buds pink, 

 the tube 4 mm. long, pinkish ; limb 6-8 mm. broad ; appendages small, one-sixth as long as the 

 corolla-lobes ; stamens inserted on upper part of corolla-tube ; nutlet 5 mm. long, broad-ovoid, 

 dorsal face hispidulous, covered all over with long barbed prickles. 



Moist places, Canadian Zone; Sierra Nevada from Plumas County to Fresno County, California. Type 

 locality: head waters of Carson River, Alpine County. July-Aug. 



18. Hackelia Sharsmithii I. M. Johnston. Sharsmith's Stickseed. Fig. 4219. 



Hackelia Sharsmithii I. M. Johnston, Journ. Arnold Arb. 20: 398. 1939. 

 Lappula Sharsmithii Jepson & Bailey in Jepson, Fl. Calif. 3: 308. 1943. 



Stems usually several from the branched crown of the stout perennial root, slender, 1-3 dm. 

 high, strigose with strongly reflexed hairs. Leaves green, rather thinly reflexed-strigose, the 

 lower oval to oblong-lanceolate, 2.5-7 cm. long, decurrent on the petiole of about the same 

 length or longer, the upper clasping, ovate to ovate-lanceolate, 2-3.5 cm. long, acute at apex: 

 racemes 2 or 3, terminal, 2-6 cm. long; pedicels 1-6 mm. long, recurved, strigose; calyx 2-2.5 

 mm. long, strigose; corolla blue with a yellow center, about 6 mm. broad; appendages lijnate, 

 ciliate on the margins above; nutlets ovoid, 2-2.5 mm. long, the marginal glochidiate prickles 

 subulate, often bluish at tip, distinct or united at base to form a narrow wing, dorsal prickles 1 

 to few, short. 



Shaded spots in shelter of rocks, Arctic- Alpine Zone; Mount Whitney region in Tulare and Inyo Counties, 

 California. Type locality: Lone Pine Canyon, altitude 11,000 feet, Inyo County, California. Aug.-Sept. 



