BORAGE FAMILY 551 



inflorescence of slender lax racemes, many- flowered, pedicels slender, 6-9 mm. in fruit; calyx 

 1.5-2 mm. long; corolla blue or whitish, about 3 mm. broad, appendages broader than long, 

 scarcely retuse, protuberances not closing throat, very finely papillose ; face of_ nutlet narrowly 

 ovate, scarcely 2 mm. long, muricate and sparsely hispidulus, occasionally with 2 or 3 short 

 prickles, the marginal glochidiate prickles 10-12 on each side, broadened at base, some short but 

 approximately equal length, the longest equaling the width of the nutlet. 



Wooded slopes and thickets, Transition Zones; British Columbia south to Okanogan County, Washington, 

 and east to Ontario and Michigan and Iowa. Type locality: not given. June-July. 



2. Hackelia floribunda (Lehm.) I. M. Johnston. Many-flowered 



Stickseed. Fig. 4203. 



Echinospermutn fioribundum Lchtn. Stirp. Pug. 2: 24. 1830. 



Lappula floribunda Greene, Pittonia 2: 182. 1891. 



Hackelia floribunda I. M. Johnston, Contr. Gray Herb. No. 68: 46. 1943. 



Stem erect, stout from a short-lived perennial root, 5-12 dm. high, the rough pubescence 

 deflexed, mixed with some spreading hairs. Leaves oblanceolate to lanceolate, hirsutulous- 

 appressed, basal leaves petiolate with spreading hairs, the stem-leaves sessile above; racemes of 

 the inflorescence many, rather strict, densely flowered, pedicels short, about 5-7 mm. long m 

 fruit ; corolla blue, 5-7 mm. broad, appendages small, obscurely papillate, not closing the throat ; 

 face of nutlet with a median ridge, muriculate, hirsutulous, without short glochidiate prickles, 

 the marginal spines much flattened at base, distinct or somewhat confluent, 4-6 on each side, 

 mostly exceeding in width the face of the nutlet. 



Brushy slopes and borders of woods, Transition and Canadian Zones; British Columbia and Washington 

 and Oregon east of the Cascade Mountains to Mono County, California; also western Ontario and Minnesota 

 to northern New Mexico and Arizona. Type locality: "Lake Pentanguishene to the Rocky Mountains. Drutnmond, 

 according to Lehmann in Hooker, Fl. Bor. Amer. 2: 84. 1838. June-July. 



3. Hackelia Jessicae (McGregor) Brand. Jessica's Stickseed. Fig. 4204. 



Lappula micrantha Eastw. Bull. Torrey Club 30: 497. 1903. Not Hackelia micrantha Opiz. 



Lappula Jessicae McGregor, Bull. Torrey Club 37: 262. 1910. 



Hackelia Eastwoodiac I. M. Johnston, Contr. Gray Herb. No. 68: 47. 1923. 



Hackelia Jessicae Brand, Pflanzenreich i'^^: 132. 1931. 



Lappula floribunda var. Jessicae Jepson & Hoover in Jepson, Fl. Calif. 3: 307. 1943. 



Stems erect or ascending, from a stout root, sparsely to rather densely villous-hirsute. 

 Basal leaves 8-15 cm. long, the blades oblanceolate, 15-20 mm. wide, narrowed to a wmged 

 petiole of about equal length; upper stem-leaves sessile, lanceolate, acute, the reduced ones 

 subtending the lower racemes often ovate-lanceolate; racemes several in an open panicle; 

 pedicels slender, at length recurved-reflexed, 5-10 mm. long ; calyx-lobes oblong to oblong- 

 lanceolate, 2-3 mm. long; corolla small, pale blue, 3.5-5 mm. broad, tube often whitish, 1.5-2 

 mm. long, lobes oblong-obovate, crests yellowish, rounded, puberulent; nutlets 4-6 mm. long, 

 marginal prickles broadly dilated at base, about 10, distinct, often with a shorter one in between, 

 dorsal face broadly ovate, usually flattened with an indistinct median ridge, puberulent and in 

 age more or less muriculate, usually with 1 or more short barbed prickles near the center. 



Usually on moist banks or slopes, Canadian Zone; British Columbia, Washington and Idaho south to 

 North Coast Ranges and the Sierra Nevada, California, and western Nevada. A variable species apparently 

 intergrading with H. Cusickii, a species to which it is allied. Type locality: Halfmoon Lake, Eldorado County, 

 California. June-Aug. 



4. Hackelia Cusickii (Piper) Brand. Cusick's Stickseed. Fig. 4205. 



Lappula Cusickii Piper, Bull. Torrey Club 29: 542. 1902. 



Lappula arida var. Cusickii Nels. & Macbr. Bot. Gaz 61: 41. 1916. 



Hackelia arida var. Cusickii I. M. Johnston, Contr. Gray Herb. No. 68: 48. 1923. 



Hackelia Cusickii Brand, Pflanzenreich A^'-'^: 131. 1931. 



Perennial, stems erect or ascending, 15-30 cm. high, crown of the woody caudex usually 

 thickly clothed with old leaf-bases, whole plant bluish-canescent with rather densely appressed 

 puberulence. Lower leaves many, linear-lanceolate or narrowly oblanceolate, 4-10 mm. wide, 

 acute, more or less hirsute-ciliate on the elongated petioles, upper stem-leaves sessile and much 

 shorter, ciliate on the margin of the blade at base ; inflorescence racemose-corymbose, with about 

 5 branches; calyx-lobes finear. acute, 2 mm. long; corolla blue, 5-6 mm. broad, appendages 

 short-pilose, about as long as broad, emarginate ; nutlets narrowly ovate, 4-5 mm. long, their 

 marginal prickles a little shorter than the width of the nutlet, 3-5 broad-based long prickles on 

 each side with much shorter slender prickles in between, all somewhat united at base; dorsal 

 face of nutlet with a faint central ridge, densely muriculate and with a few irregularly dispersed 

 short slender barbed prickles. 



Drv slopes, often among junipers, mainly Arid Transition Zone; eastern Oregon from Gilliam County 

 to Klamath and Malheur Counties, and to Modoc County, California. Type locality: "Logan Mountains, 

 eastern Oregon, 6500 feet altitude, in the shelter of juniper." May-July. 



Hackelia saxatilis (Piper) Brand, Pflanzenreich 42^'2. 133. 1931. {Lappula saxatilis Piper, Bull. Torrey 

 Club 29: 541. 1902.) Stems rather slender, 2-3 dm. high, herbage thinly cinereous throughout with a fine 

 soft spreading and reflexed pubescence and occasional longer villous hairs intermixed, especially on the petioles 

 of the lower leaves and on the margins of the upper ones. Lower leaves oblanceolate, acute, with petioles 

 about as long as the blades; upper leaves linear-lanceolate, sessile; branches of the spreading inflorescence 6-10- 

 flowered; corolla blue, rotate, about 7 mm. broad; appendages pubescent, broader than long; nutlets 3-4 mm. 



