480 HYDROPHYLLACEAE 



Moist woods and canyons, Upper Sonoran and Arid Transition Zones; Coast Ranges, Sierra Nevada foot- 

 hills, and Santa Barbara Islands of California. Type locality: California. March-June. 



Pholistoma auritum var. arizonicum (M. E. Jones) Constance, Bull. Torrey Club 66: 348. 1939. (Nemo- 

 phila arizonica M. E. Jones, Contr. West. Bot. No. 12: 50. 1908.) Smaller and more slender, less prickly through- 

 out; leaf-divisions fewer, blunt; corolla usually less than 1 cm. broad, about equaling the calyx. Mountains of 

 southeastern California to Arizona. Type locality: "Chimihuevis Mountains, Arizona." 



2. Pholistoma racemosum (Nutt.) Constance. San Diego Fiesta-flower. 



Fig. 4047. 



Nemophila racemosa Nutt. ex A. Gray, Proc. Amer. Acad. 10: 315. 1875. 

 Nemophila erodiifolia Millsp. in Millsp. & Nutt. Field Mus. Bot. Ser. 5: 205. 1923. 

 Fholistoma racemosum Constance, Bull. Torrey Club 66: 349. 1939. 

 Ellisia racemosa Jepson, Fl. Calif. 3: 237. 1943. 



Stems Z-6 dm. long, sparingly hispidulous. Lower leaves ovate or deltoid-ovate, 4-10 cm. 

 long, 2-6 cm. broad, obtuse at apex, subcordate or truncate at base, the divisions 5-9, oblong 

 or ovate, more or less hispidulous on both surfaces, more densely so beneath, the petioles 

 narrowly winged, but not auriculate; flowers solitary, or usually 2-6 in cymes; calyx-lobes 

 linear- or ovate-lanceolate, 2-3 mm. long, 1-1.5 mm. broad, the auricles 1-1.5 mm. long; 

 corolla 0.6-1 cm. broad, white or blue, the lobes obovate ; appendages triangular but usually 

 narrow or obsolete; style about 2.5 mm. long; capsule 5-8 mm. in diameter, enclosed by the 

 calyx; seeds usually 4-6, sometimes 1, 1-2 mm. in diameter. 



Moist, shaded places, Sonoran Zones; San Diego and Lower California coast, and Santa Cruz, San Cle- 

 mente, Santa Catalina, and Mexican islands. Type locality: San Diego. Feb.-May. 



3. Pholistoma membranaceum (Benth.) Constance. White Fiesta-flower. 



Fig. 4048. 



Ellisia membranacea Benth. Trans. Linn. Soc. 17:274. 1835. 

 Ellisia membranacea var. hastifolia Brand, Pflanzenreich 4^^: 38. 1913. 

 Pholistoma membranaceum Constance, Bull. Torrey Club 66: 350. 1939. 



Stems 0.5-6 dm. long, glabrous. Lower leaves oblong to oval, 3-12 cm. long, 1-5 cm. broad, 

 obtuse at apex, subcordate or truncate at base, the divisions 5-11, oblong, obtuse, rather distant 

 and entire or with a single tooth, sparsely hispidulous on both surfaces, the petioles more or less 

 winged but not auriculate; flowers 2-10 in cymes, rarely solitary; calyx-lobes oval, 2-3 mm. long, 

 1-2 mm. broad, auricles none; corolla 0.4-1 cm. broad, white and often with a lanceolate purple 

 spot on each lobe, the lobes oval ; appendages triangular, very small ; style 1 . 5-2 mm. long ; 

 capsule 2-4 mm. in diameter, the fruiting calyx stellate-spreading ; seeds 1-2, 2-3 mm. in diameter. 



Moist shade of oaks or bushes, Sonoran Zones; Coast Ranges and Sierra Nevada foothills of central California 

 to the desert ranges of Inyo and San Bernardino Counties to southern Nevada and Lower California, absent from 

 the islands and usually from the immediate coast. Type locality: California. March— June. 



3. NEMOPHILA Nutt. ex Barton, Fl. N. Amer. 2: 71. 1822; Journ. Acad. 

 Phila. 2: 179. 1822. Nomen conservandum. 



Delicate, weak, usually branched, hispid or glabrous annual herbs from slender tap- 

 roots. Leaves all opposite, all alternate, or the lower opposite and the upper alternate, 

 variously toothed, lobed, or pinnately divided, petiolate. Flowers solitary in the upper 

 axils or opposite the leaves, pedicellate. Calyx divided nearly to the base, the sinuses 

 armed with a sepaloid auricle, or the auricles obsolete. Corolla white or blue, plain or 

 variously marked, deciduous, campanulate to rotate, divided one-third to two-thirds, 

 longer or shorter than the calyx. Stamens included, equal and equally inserted on the 

 corolla ; appendages a pair to each filament, adnate or partially free, ciliate to glabrous, 

 or reduced to hairy lines or obsolete. Style shallowly to deeply bifid. Mature capsule 

 unilocular, ovoid or globose, loculicidally dehiscent. Ovules two to several on the front 

 of each of the two large parietal placentae. Seeds 1-20, ovoid, corrugate-tuberculate or 

 smooth, regularly or irregularly pitted or without pits, yellow, red, brown, or black, pro- 

 vided with a shallow or papillae-form group of colorless cells (cucullus), which is con- 

 spicuous or reduced, deciduous or persistent. [Name Greek, meaning grove and to love.] 



A genus of 11 species, chiefly of the western United States, extending to neighboring British Coluimbia and 

 Lower California, and with 2 species in the southeastern states. Type species, Nemophila phacelioides Nutt. 



Stems minutely recurved-prickly; leaves all alternate; corolla shorter than the calyx; seed globose, solitary; cu- 

 cullus persistent. 1. A', breviflora. 



Stems variously hispid or glabrate, never prickly; leaves all opposite or the upper alternate; corolla equaling or 

 exceeding the calyx; seeds more or less ovoid, 2 to several; cucullus deciduous. 



Capsule exceeded by the fruiting calyx; seeds pitted in rows; cucullus reduced. 2. N. Kirtleyi. 



Capsule equaling or exceeding the fruiting calyx; seeds irregularly pitted or without pits; cucullus con- 

 spicuous. 



Corolla 1 cm. or more broad; leaves all opposite. 



Seeds corrugate-tuberculate; lower surfaces of leaves pale, bearing stomata; corolla white or blue 



and variously marked, but the lobes not purple-blotched. 4. A^. Menzicsii. 



Seeds smooth or pitted; both surfaces of leaves alike, bearing stomata; corolla white and venose, but 



with a purple blotch at the tip of each lobe. 6. N. maculata. 



