PHLOX FAMILY 455 



axils or in few-flowered cymes. 5- or sometimes 6-merous ; calyx deeply cleft into subequal linear 

 acerose lobes, sinuses one-half to two-thirds filled with a broad hyahne membrane formmg a 

 definite pseudotube : corolla salverform, pink, rarely white, tube and throat contmuous, 10-15 mm. 

 long lobes twisted in bud, rotately spreading in anthesis, limb 20-30 mm. broad ; stamens inserted 

 near' the middle of tube, filaments about 1 mm. long ; stigma included ; capsule ellipsoid, locules 

 several-seeded. 



Coastal mesas and canyons and in the chaparral Upper Sonoran Zone; South Coast Ranges mCahforma. ■ 

 from San Luis Obispo County to the Santa Monica Mountains. Type locality: California. Collected by Douglas, 

 probably on his trip from Monterey to Santa Barbara. March-June. 



Leptodactylon californicum subsp. glandulosum (Eastw.) H. L. Mason. (^Gilia calif ornica var. dla^'duhf 

 Eastw Bot Gaz 37:447. \904: Leptodact\!on calif ormcum glandulosum Ahrams, BuU.N.Y . Bot. (jard.t . 'iJ:^. 

 1910 L californicum f. nlandulosum Wherry, Amer. Midi. Nat. 34:383. 1945.) Herbage very pubescent; 

 leaves often arachnoid-glandular. Mountains of southern California. Type locality: Pasadena. California. 



2. Leptodactylon pungens (Terr.) Rydb. Granite Gilia. Fig. 4009. 



Cantua pungcns Torr. Ann. Lye. N.Y. 2: 221. 1828. 



Batanthes pungens Raf. Atl. Journ. 145. 1832. 



Acgochloa Torrcyi G. Don, Gen. Hist. PI. 4: 246. 1838. 



GiVio />i(Mf7en.s Benth. in A. DC. Prod. 9:316. 1845. 



Navarretia pungens Kuntze, Rev. Gen. PI. 2: 433. 1891. 



Leptodactylon pungens Rydb. Fl. Colo. 279. 1906. 



Gilia pungens subsp. cu-pungens Brand, Pflanzenreich 4-"': 126. 1907. 



Leptodactylon Haccliae M. E. Peck, Proc. Biol. Soc. Wash. 49: 111. 1936. 



Leptodactylon pungens subsp. eupungens Wherry, Amer. Midi. Nat. 34: 383. 1945. 



Leptodactylon pungens subsp. Hookeri f. Hazcliae Wherry, loc. cit. 



Shrubs 1-8 dm. high, erect, spreading, internodes subequaling leaves, rarely longer, herbage 

 densely glandular-villous. Leaves alternate, occasionally subopposite or sometimes the lower 

 opposite, sessile, palmately cleft into 3-7 unequal, linear, pungent or rigidly acerose lobes with 

 others fascicled in the axils ; flowers sessile to subsessile in terminal or axillary glomerules or 

 sometimes solitary in upper axils ; calyx deeply cleft into unequal, linear, acerose lobes, sinuses 

 about two-thirds filled with a hyaline membrane forming an evident pseudotube; corolla nar- 

 rowly funnelform, 1-2 cm. long, white, yellow, lilac, or pink, often sordid and with a brownish 

 purple pigment on back of lobes, throat very narrow, almost continuous with tube, lobes sub- 

 rotately spreading, very sensitive to fluctuations in light intensity in opening and closing; sta- 

 mens inserted on throat, included, filaments subequal to shorter than anthers, glabrous; pistil 

 about one-third as long as corolla-tube ; capsule cylindric, locules many-seeded. 



Dry places and rocky ridges, Arid Transition Zone; British Columbia, Washington, and eastern Oregon, 

 south through the Sierra Nevada to the mountains of Lower California, and east to the Rocky Mountains and 

 New Mexico. The subspecies of Leptodactylon pungens are not too well defined. Type locality: "valley of the 

 Loupe Fork," Nevada. May-Aug. 



Leptodactylon pungens subsp. pulchriflora (Brand) H. L. Mason. (Gilia pungens var. squarrosa A. Gray, 

 Proc Amer Acad 8: 268. 1870; Cantua pungens var. squarrosa Howell, Fl. N.W. Amer. 453. 1901; Leptodac- 

 tylon lilacinum Greene ex C. F. Baker, West. Amer. PI. 1 : 18. 1902. _ [Nomen nudum] ; L. ^patens HeUer, Muh- 

 lenbergia 1:146. 1906; G. pungens subsp. eu-pungens var. devesttta Brand, Pflanzenreich 4^: 128. lyu/; 

 G lilacina Brand, loc. cit.; G. pungens subsp. pulchriflora Brand, Ann. Conserv. & Jard. Bot. Geneve IS: 333. 

 1913- L pungens squarrosum Tidestrom, Proc. Biol. Soc. Wash. 48:42. 1935; L. pungens var. suhflavidum 

 Jepso'n Fl. Calif. 3: 143. 1943; L. lilacinum f. pulchriflorum Wherry, Amer. Midi. Nat. 34: 384. 1945.) Lateral 

 leaf-segments rarely less than half the length of the middle lobe, yet very unequal, often much branched; corolla- 

 tube often long-exserted. By far the common subspecies in the Pacific States. Washington and Oregon and the 

 mountains of western Great Basin through the Sierra Nevada to the mountains of southern California. Type 

 locality: Farewell Gap, Tulare County, California. 



Leptodactylon pungens subsp. Hookeri (Dougl.) Wherry, Amer. Midi. Nat. 34: 383. 1945. (Phlox 

 Hookeri Dougl. ex Hook. Fl. Bor. Amer. 2: 7i. 1838; Gilia Hookeri Benth. in A. DC. Prod. 9: 316. 1845; Lepto- 

 dactylon Hookeri Nutt. journ. Acad. Phila. 1: 157. 1848; G. pungens var. Hookeri A. Gray, Proc. Amer. Acad. 

 8- 268 1870; Cantua pungens var. Hookeri Howell, Fl. N.W. Amer. 453. 1901; G. pungens subsp. eu-pungens 

 var. Hookeri Brand, Pflanzenreich 4==*°: 126. 1907; L. brevifolium Rydb. Bull. Torrey Club 40:474 1913; 

 L. pungens var. Hookeri Jepson, Man. Fl. PI. Calif. 807. 1925; L. pungens var. shastense Jepson F . Calif. 

 3: 143. 1943; L. pungens subsp. brevifolium Wherry, Amer. Midi. Nat. 34: 383. 1945; L. Itlactnum f. shastense 

 Wherry, op. cit. 384.) Leaves very short, often less than 5 mm., opposite in north of range, subopposite to alter- 

 nate in south of range; stamens midway on throat; branching usually virgate. Eastern Washington southward to 

 Arizona. Type locality: "near the narrows of the Oakangan and Priest's Rapid of the Columbia. 



Leptodactylon pungens subsp. Hallii (Parish) H. L. Mason. (Gilia Hallii Parish, Erythea 7: 94. 1899; 

 G. tenuiloha Parish, op. cit. 95; G. punaens var. Hallii Milliken. Univ. Calif. Pub. Bot. 2: 42. 1904; G. pungens 

 var. tenuiloha Milliken, op. cit. 43; Leptodactylon Hallii Heller, Muhlenbergia 1: 146. 1906; L. tcnutlobum 

 Heller loc cit.; G. pungens subsp. eu-punqens var. tenuiloha Brand, Pflanzenreich A^^: 128. 1907; G. pungens 

 subsp. Hallii Brand, loc. cit.; L. pungens var. tenuilobum Jepson, Man. Fl. PI. Calif. 807. 1925; L. pungens var. 

 Hallii, loc. cit.) Middle leaf-segment divaricate, spinose, lateral segments rarely over one-third as long, occasionally 

 quite obsolete; stems often virgate; texture of foliage quite variable. Eastern Washington and Idaho south through 

 the Great Basin to the mountains of southern California. Type locality: Coyote Canyon, El Toro Mountain, 

 Riverside County, California. 



3. Leptodactylon Jaegeri (Munz) Wherry. San Jacinto Phlox. Fig. 4010. 



Gilia Jaegeri Munz, Bull. S. Calif. Acad. 31 : 68. 1932. 



Leptodactylon pungens var. Jaegeri McMinn, Man. Calif. Shrubs 450. 1939. 

 Leptodactylon pungens Jepson, Fl. Calif. 3: 142, in part. 1943. 

 Leptodactylon Jaegeri Wherry, Amer. Midi. Nat. 34: 384. 1945. 



Cespitose perennials with woody base, 3-10 cm. high. Leaves chiefly opposite, simple or 

 3-cleft into linear, flat, strongly venose lobes from the end of a conspicuous petiole; flowers 

 solitary in the upper leaf-axils; calyx narrowly cylindric, 8-10 mm. long; sepals 6, subequal, 

 sinuses of the lobes nearly filled with a narrow hyaline membrane; corolla funnelform, 25-30 

 mm. long with 6 (rarely 5) petals, tube 2 times the calyx ; stamens 6, inserted on base of throat, 



