574 BORAGINACEAE 



mature nutlets varying from 1-4 but usually 3, orbicular-ovoid, abruptly constricted into an 

 acute beak-like apex, median dorsal keel and lateral ones usually prominent, transverse rugae 

 4 or 5, the intervals rectangular, minutely papillate; scar annular, median at the base of the 

 narrow ventral keel. 



Grassy fields and hillsides, Upper Sonoran Zone; Klickitat County, Washington, and Wasco County, Oregon, 

 southward mostly west of the Cascades to San Diego County and east to the Providence Mountains, California. 

 Type locality: California. Collected by Douglas. March-May. 



Plagiobothrys myosotoides (Lehm.) Brand, Pflanzenreich ^^': 108. 1931. (Lithospermiim tinctorium Ruiz 

 & Pavon, Fl. Peruv. 2:4. pi. 114. fig. 6. 1799. Not L. 1753; L. myosotoides Lehm. Asperif. 319. 1818; 

 Plagiobothrys tinctorius A. Gray, Proc. Amer. Acad. 20: 283. 1885.) In habit much like P. Torreyi but the 

 sculpturingof the nutlets more like that of P. tenellus; strongly keeled dorsally and also with ridges separated 

 by broad intervals that are sometimes papillate, not shining. A Chilean species of which only two collections 

 have been reported from California: ridge between Isabel Valley and Arroyo Bayo, Mount Hamilton Range, and 

 Big Sandy Valley, Black Mountain, Fresno County. 



6. Plagiobothrys uncinatus J. T. Howell. Hooked Popcorn Flower. Fig. 4257. 



Plagiobothrys uncinatus J. T. Howell, Leaflets West. Bot. 2: 255. 1940. 



Stems usually several from the base and bushy-branched, 8-30 cm. high, reddish, thinly 

 hirsute with spreading hairs, without finer pubescence. Basal leaves linear-oblong, 2-2.5 cm., 

 long including the petiole, 3-4 mm. wide, hispidulous, cauline leaves oblong-ovate to narrowly 

 ovate, 3-10 mm. long; flowers scattered along the stems and solitary in the leaf-axils, also in 

 small terminal clusters, subsessile ; calyx divided almost to base, densely uncinate-bristly, lobes 

 about 1 mm. long in anthesis, 2-2.5 mm. in fruit; corolla 1.5 mm. long, about 1 mm. broad; nut- 

 lets 1-1.3 mm. long, somewhat quadrate, rounded at base, abruptly pointed at apex, the dorsal 

 side slightly keeled toward the apex, transversely rugulose and tuberculate, the ventral side 

 narrowly keeled above the middle. 



Slopes of canyons, Upper Sonoran Zone; Gabilan and Santa Lucia Mountains, California. Type locality: 

 Santa Lucia Camp, Santa Lucia Mountains, Monterey County. April-May. 



7. Plagiobothrys campestris Greene, Fulvous Popcorn Flower. Fig. 4258. 



Plagiobothrys calif ornicus Greene, Vittonia 2: 231. 1892. Not Greene, Bull. Calif . Acad. 2 : 407. 1887. 



Plagiobothrys campestris Greene, Pittonia 2: 282. 1892. 



Plagiobothrys rufescens var. campestris Jepson, FI. W. Mid. Calif. 446. 1901. 



Plagiobothrys fulvus var. campestris I. M. Johnston, Contr. Gray Herb. No. 68: 70. 1923. 



Stem solitary or rarely 2 or more from the base, 3-6 dm. high or the branches sometimes 

 spreading and more or less decumbent, villous-hirsute with spreading hairs, and sparsely to- 

 mentose. Leaves hirsute with appressed hairs on both surfaces and spreading ones on the 

 margins, the basal spatulate-oblanceolate, the cauline sessile and linear, the lower 3-5 cm. 

 long, the upper gradually shorter ; spicate racemes loosely flowered, often 3-4 dm. long ; calyx 

 often fulvous when young, the lobes lanceolate, about 5 mm. long and erect in fruit; corolla 

 3-^ mm. broad; nutlets 2.5-3 mm. long, triangular-ovoid, short-acuminate, dorsal side thin- 

 keeled, the median keel extending over the apex, transverse rugae usually several or sometimes 

 nearly obsolete ; scar annular. 



Open valleys and foothills, Upper Sonoran Zone; Umpqua Valley, western Oregon, southward west of the 

 Cascade-Sierra Nevada Divide, to Santa Clara County in the Coast Ranges and to the foothills of the Sierra 

 Nevada, Fresno County, California. Type locality: "Interior California." March-May. 



8. Plagiobothrys infectivus I. M. Johnston. Dye Popcorn Flower. Fig. 4259. 



Plagiobothrys infectivus I. M. Johnston, Journ. Arnold Arb. 20: 380. 1939. 



Stem solitary and erect or several from the base and spreading, 2-4 dm. long, villous-hirsute 

 with spreading hairs or somewhat retrorse; root, lower part of stem, midvein and margin of 

 leaves purple-stained. Leaves linear, 2-8 cm. long, nearly glabrous beneath, appressed-hirsute 

 above; spikes 10-20 cm. long, loosely flowered, leafy-bracteate throughout; calyx cleft to the 

 base into broadly lanceolate acuminate lobes, 5-7 mm. long, rusty-hirsute when young, mid- 

 vein purple-stained ; corolla slightly exserted, 3 mm. broad, often rose-colored in drying ; mature 

 nutlets usually 4, broadly ovoid, 3-3.5 mm. long, abruptly constricted into a short acute apex; 

 median keel thin, prominent at least on the acute beak, lateral keels distinct on the beak but 

 sometimes obscure on the body, ventral keel prominent above the median scar, this raised 

 and annular, dorsal surface tuberculate, with few or no rugae, the ventral surface with several 

 irregular rugae. 



Usually in adobe soils, on open hills. Upper Sonoran Zone; Inner Coast Ranges especially on the eastern side, 

 from Colusa County to San Luis Obispo County, California. Type locality: Hospital Canyon, western San 

 Joaquin County. March-May. 



9. Plagiobothrys canescens Benth. Valley Popcorn Flower. Fig. 4260. 



Plagiobothrys canescens Benth. PI. Hartw. 326. 1849. 

 Eritrichium canescens A. Gray, Proc. Amer. Acad. 10: 53. 1874. 

 Plagiobothrys microcarpa Greene, Pittonia 1: 21. 1887. 



Stems usually with several branches from or near the base, ascending or more commonly 

 decumbent and widely spreading, villous-tomentose throughout. Leaves linear or the basal 

 linear-oblanceolate, 1.5-5 cm. long; spikes elongated and loosely flowered in age, bracteate; 

 fruiting calyx 4-6 mm. long, densely villous-hirsute, cleft to below the middle, the lobes erect 

 or rarely curved over the nutlets, broadly lanceolate, somewhat acuminate at apex; corolla 

 about 3 mm. broad ; nutlets usually 4, orbicular-ovoid, constricted above into a short beak-like 



