HEATH FAMILY 323 



after fire, the young branchlets, petioles and leaves glandular-hairy. Leaves ovate to ovate- 

 lanceolate, 3-5 cm. long, acute or obtuse at apex, obtuse or rounded at base, dull green and 

 similar on both sides ; bracts f oliaceous ; pedicels viscid-tomentose ; corolla white, 7-8 mm. long ; 

 fruit depressed-globose, viscid-puberulent. 



Dry rocky slopes and ridges, Humid Transition and Upper Sonoran Zones; Coos Bay, Oregon, south in 

 the Coast Ranges to Orange County, California. Type locality: Mount Tamalpais, Marin County, California. 

 Feb.-April. Crown Manzanita. 



This species is extremely variable and exhaustive experimental and genetic studies as well as further 

 field studies are necessary to determine the specific limits and the range of variability. Dr. J. E. Adams, long 

 a student of the genus in the field and herbarium, has proposed the following varieties: 



Leaves light green or pallid. 



Branchlets not glandular. 



Inflorescence and ovary not glandular; leaves light yellowish green. var. Howellii. 



Inflorescence and ovary glandular. 



Branchlets canescently tomentulose; leaves with a light bloom. var. Cushingiana. 



Branchlets long white-hairy. 



Inflorescence many-flowered and spreading. var. Campbelliae. 



Inflorescence few-flowered and compact. var. mollis. 



Branchlets glandular-hairy, as in the typical species; leaves very pallid, scabrous. 



var. sacaensis. 



Leaves dark green. var. crassifolia. 



Arctostaphylos glandulosa var. Howellii (Eastw.) J. E. Adams ex McMinn, 111. Man. Calif. Shrubs. 

 417. 1939. (Arctostaphylos Hoivcllii Ecistw. Leaflets West. Bot. 1 : 123. 1934.) Erect shrub, 1-2 m. high, with 

 spreading branches, old stems polished dark reddish brown, young branchlets finely tomentose. Leaves yellow- 

 ish gray-green, oblong-elliptic to suborbicular, 3-5 cm. long, sparsely tomentose on both sides; branches of 

 the panicle glandular-pubescent; bracts linear or triangular-subulate; fruit depressed-globose, sparsely tomen- 

 tose and glandular-viscid. Central Monterey County, California. Type locality: Arroyo Seco Canyon, Santa 

 Lucia Mountains. 



Arctostaphylos glandulosa var. Cushingiana (Eastw.) J. E. Adams ex McMinn, loc. cit. (Arctostaphylos 

 Cushingiana Eastw. Leaflets West. Bot. 1:75. 1933.) Low densely branched shrub, 8-12 dm. high, old 

 trunks dark reddish purple, branchlets tomentose and with scattering short hairs, somewhat viscid-glandular. 

 Leaves ovate or elliptic to lanceolate, 2-4 cm. long, pallid gray-green, minutely tomentulose or soon glabrate; 

 bracts f oliaceous, glandular-tomentose and ciliate on the margins; pedicels densely white-tomentose; top of ovary 

 sparingly white-pubescent; fruit depressed-globose, glabrous. Dry mountain slopes; Napa, Sonoma and Marin 

 Cfounties, California. Type locality: south side of Mount Tamalpais, Marin County. 



Arctostaphylos glandulosa var. Campbelliae (Eastw.) J. E. Adams ex McMinn. loc. cit. (Arctostaphylos 

 Campbelliae Eastw. Leaflets West. Bot. 1: 74. 1933.) Low shrub, with the branchlets rather densely puberulent 

 or tomentulose with intermingling coarse white spreading hairs. Leaves yellowish green, short-pubescent 

 below, less so or glabrous above, acute at apex, obtuse at base; bracts foliaceous, lanceolate, the upper ones 

 becoming broadly deltoid; ovary white-hairy, not glandular; fruit sparsely hairy. Dry ridges. Mount Hamilton, 

 California. A local variation apparently restricted to the immediate vicinity of the mountain. 



Arctostaphylos glandulosa var. mollis J. E. Adams, Journ. E. Mitchell Sci. Soc. 56: 50. 1940. Erect 

 shrub, 1-1.5 m. high, divaricately much-branched; young branchlets pubescent and white-hairy, not glandular. 

 Leaves pallid green, glabrescent, elliptic to ovate, acute at apex, acute or obtuse at base, 2-4 cm. long; 

 panicles small, compact, few-flowered; bracts foliaceous, hirsute-ciliate; pedicels pubescent or somewhat 

 glandular; ovary densely white-villous, not glandular. Santa Ynez Mountains, Santa Barbara County to the 

 San Gabriel Mountains, Los Angeles County, California. Type locality: La Curabre Peak, Santa Barbara County. 



Arctostaphylos glandulosa var. zacaensis (Eastw.) J. E. Adams ex McMinn, Til. Man. Calif. Shrubs 

 417. 1939. (Arctostaphylos zacaensis Eastw. Leaflets West Bot. 1: 79. 1933.) Much like the typical species in 

 habit and its glandular pubescence, but leaves very pale grayish green, pubescent and scabrous and slightly 

 glandular-hairy at base, 4-6 cm. long, 2-4 cm. wide, obtuse or acute and apiculate at apex, obtuse at base. 

 Rocky slopes in the vicinity of Zaca Lake, Santa Barbara County, California. Type Locality: Zaca Lake. 



Arctostaphylos glandulosa var. crassifolia Jepson, Madroiio 1 : 86. 1922. Erect shrub, about 1 m. 

 high; young branchlets tomentulose and with or without a few scattering hairs, not glandular. Leaves dark 

 green, dull or a little shiny, broadly elliptic to oval, acute to rounded at apex, mostly obtuse at base, 2-4.5 cm. 

 long, glabrate above, tomentulose or glabrate beneath; pedicels pubescent; fruit sparsely pubescent or glabrate, 

 not glandular. Sandy soils on the mesas near the coast, Oceanside to San Diego, San Diego County, California. 

 Type locality: Del Mar, San Diego County. 



38. Arctostaphylos intricata Howell. Siskiyou Manzanita. Fig. 3731. 



Arctostaphylos intricata Howell, Fl. N.W. Amer. 416. 1901. 



Erect shrub, 1-2 m. high, irregularly and intricately branched, old stems polished and very 

 dark-colored, young branchlets tomentose and glandular-hispid or glandular-hispidulous. Leaves 

 oblong-lanceolate to oblong-ovate, 2 . 5-4 cm. long ; panicles short and dense ; bracts foliaceous ; 

 pedicels sparsely glandular-pubescent with spreading hairs ; corolla rose-colored or white, about 

 5 mm. long; ovary pubescent; fruit depressed-globose, pubescent, but not viscid. 



Mountain slopes and canyons, mainly Transition Zones; southern Coos and Douglas Counties, Oregon, 

 to Del Norte County, California. Type locality: stony hillsides, near Smith River, west of Gasquet, Del 

 Norte County, California. April-May. 



Arctostaphylos oblongifolia Howell, Fl. N.W. Amer. 416. 1901. (Arctostaphylos viscida var. ohlongi- 

 folia J. E. Adams ex. McMinn, 111. Man. Calif. Shrubs 403. 1939.) Erect shrub, 1-2 m. high, densely 

 branched from the base, the bark polished rich red-brown, branchlets cinereous. Leaves 2.5-3.5 cm. long, 

 oblong to broadly ovate, usually abruptly contracted at base to a terete petiole, pallid green and smooth; bracts 

 somewhat foliaceous, puberulent; pedicels longer than the bracts, pubescent and glandular; ovary pubescent; 

 fruit depressed-globose. Rocky hillsides. Arid Transition Zone; southern Douglas County, Oregon, to Del 

 Norte County, California. Type locality: "Rocky hillsides, eastern base of the coast mountains near Waldo, 

 Oregon." 



39. Arctostaphylos Crustacea Eastw. Brittle-leaved Manzanita. Fig. 3732. 



Arctostaphylos Crustacea Eastw. Leaflets West. Bot. 1:74. 1933. 



Erect branching shrub, 1-2 m. high, the stems enlarged at the base and stump-sprouting after 

 fire, dark reddish purple, young branchlets tomentose and more or less densely setose-bristly, 



