654 MENTHACEAE 



apex, membranous and rose-purple at least at tip, pubescent on the veins and ciliate on the mar- 

 gins; calyx 6-8 mm. long, 13-nerved; corolla pale purple, about 15 mm. long, tube well-exserted, 

 lobes very slender. 



This species, which was first discovered by Douglas near the narrows above Kettle Falls on the Columbia 

 River, Washing^ton, is one of the most variable of all our native plants. The complex as a whole ranges 

 from British Columbia to northern Lower California on the Pacific Coast, and eastward to Montana, Colorado 

 and New Mexico; Arid Transition and Canadian Zones. Many specific segregates have been described, some 

 of which are fairly well-defined geographic entities, and Epling (Ann. Mo. Bot. Card. 12: 58-70. 1925) has 

 proposed the following subspecies: 



Monardella odoratissima subsp. euodoratissima Epling, Ann. Mo. Bot. Card. 12:59. 1925. Branches 

 thinly pubescent above, hardly cinereous; leaves lanceolate, subsessile, green, appearing glabrous but sparingly 

 puberulent; bracts ovate to suborbicular, pubescent on the veins; calyx woolly-pubescent around the teeth; 

 corolla about 15 mm. long, usually pale purple and exserted. Northeastern Washington to northeastern Oregon, 

 and locally in Nevada and New Mexico. Ocassionally plants resembling this, the typical species, are found in 

 southern Oregon and in northern California. Type locality: near the narrows above Kettle Falls on the 

 Columbia River, Washington. 



Monardella odoratissima subsp. pinetorum (Heller) Epling, op. cit. 68. (Monardella pinetorutn Heller, 

 Muhlenbergia 1:36. 1904.) Branches minutely villous-pubescent, not glaucous; leaves ovate to lanceolate, 

 l.S-2.5 cm. long, abruptly narrowed to a usually margined petiole 2-8 mm. long, softly pubescent or short- 

 villous; bracts ovate, erect, equaling the calyces, short-pubescent, purplish; calyx pubescent with straight 

 spreading hairs; corolla rose-purple, the lobes distinctly tapering, the tube slightly exserted. Rocky situations. 

 Arid Transition Zone; Coast Ranges in Lake and Glenn Counties, also southern Sierra Nevada in Fresno and 

 Tulare Counties, California. Type locality: "southern slope of Mt. Sanhedrin, Lake county, above the sawmill, 

 in dry gravelly ground among pine trees." 



Monardella odoratissima subsp. parvifolia (Greene") Epling, op. cit. 69. (Monardella parvifolia Greene, 

 PI. Baker. 3: 22. 1901; M. muriculaia Greene, Pittonia 5: 84. 1902.) Branches slender, mostly 1-2 dm. higb, 

 minutely puberulent and rather sparsely muriculate, sometimes reddish; leaves lanceolate or oblong, 1-2 cm. 

 long, tapering at base to a margined petiole 1-3 cm. long, sparsely cinereous-puberulent ; heads small, 1-2 cm. 

 broad; bracts small, seldom exceeding the calyces, ovate, acute, short-pubescent on the back, purplish; calyx 

 5-6 mm. long, pubescent, sparingly villous around the teeth; corolla-tube little exceeding the calyx. Rocky 

 alpine slopes and ledges. Boreal Zones; Sierra Nevada, mostly above 8,000 feet, California, ranging east to 

 Colorado and New Mexico. Type locality : canyon of the Gunnison near Cimarron, Colorado. 



Monardella odoratissima subsp. australis (Abrams) Epling, op. cit. 70. (Monardella australis Abrams, 

 Muhlenbergia 8:34. 1912.) Branches decumbent or ascending, sparsely villous-pubescent; leaves lanceolate 

 to oblong, green or cinereous, 1-2.5 cm. long, narrowed to a short petiole 1-3 mm. long; bracts lanceolate, 

 short-acuminate, exceeding the calyces, rose-purple, puberulent; corolla about 15 mm. long, rose-colored, the 

 tube slightly exceeding the calyx, the lobes slender, slightly tapering. Mountain slopes and gravelly flats. Arid 

 Transition Zone, southern California, from the San Gabriel to the San Jacinto Mountains. Type locality: 

 Tamarack Valley, altitude 9,200 feet, San Jacinto Mountains. 



Monardella odoratissima subsp. discolor (Greene) Epling, op. cit. 60. (Monardella discolor Greene, 

 Pittonia 2:24. 1889.) Branches pubescent above, scurfy and cinereous; leaves ovate-lanceolate, subsessile, 

 hoary to cinereous with a dense minute tomentum; bracts woolly-pubescent or pubescent; calyx woolly-pubescent. 

 Eastern base of the Cascade Mountains from Chelan County, Washington, to The Dalles and Mount Hood 

 region, Oregon. Type locality: gravelly banks of the Yakima near Cle Elum, Washington. 



Monardella odoratissima subsp. glaiica (Greene) Epling, op. cit. 62. (Monardella glauca Greene, 

 Pittonia 4:321. 1901; M. modocensis Greene, loc. cit.; M. rubella Greene, Pittonia 5:84. 1902.) Branches 

 puberulent appearing glaucous and often purple; leaves ovate-lanceolate to oblong, 2-4 cm. long, narrowed at 

 base to a distinct petiole 1-5 cm. long, often glaucous; outer bracts ovate, the inner usually oblong, puberulent 

 on the back, purplish; calyx pubescent, hirsute around the teeth; corolla reddish purple, the tube usually well- 

 exserted. Eastern slopes of the Ca.scade Moiintains, southern Oregon, to the Mount Shasta region, California, 

 and northwestern Nevada. Type locality: "Deserts of eastern Oregon." 



Monardella odoratissima subsp. pallida (Heller) Epling, op. cit. 66. (Monardella pallida Heller, 

 Muhlenbergia 1:36. 1904; Madronella pallida Heller, op. cit. 1:138. 1906; Monardella odoratissima var. 

 ovata (Greene) Jepson, Man. Fl. PI. Calif. 882, in part. 1925.) Branches scurfy-pubescent and cinereous, pale, 

 never purplish; leaves lanceolate-oblong, 2-3 cm. long, rather abruptly narrowed to a usually margined petiole 

 2-8 mm. long, minutely cinereous-puberulent; bracts seldom exceeding the calyces, short, purplish, often re- 

 curved; calyx woolly, usually densely so throughout, the heads appearing very compact; corolla pale, usually 

 whitish, the tube little or not at all exserted. This is the most common Monardella in the Sierra Nevada, 

 California. Arid Transition and Canadian Zones. Type locality: foot of the ridge on south side of Donner 

 Lake, Nevada County, California. 



11. Monardella Robisonii Epling. Robison's Monardella. Fig. 4441. 



Monardella Robisonii Epling in Munz, Man. S. Calif. 451, 600. 1935. 



Perennial with branching woody caudex, leaf-bearing branches 4-5 dm. high, simple or 

 branched, cinereous-hirtellous. Leaves ovate-lanceolate to narrowly ovate-elliptic, 6-15 mm. long, 

 cinereous-hirtellous especially on the veins ; petioles 1-3 mm. long ; bracts membranaceous, pallid, 

 ovate, 8-10 mm. long, hirtellous ; calyx 7-8 mm, long, the teeth narrowly deltoid, 1-1.5 mm. long ; 

 corolla pale, the tube slightly exserted, the lobes about 2 mm. long. 



Rocky desert slopes. Upper Sonoran Zone; Little San Bernardino Mountains, San Bernardino County, 

 California. Type locality: "Keyes Ranch, Little San Bernardino Mts., among boulders and in crevices." June. 



12. Monardella linoides A. Gray. Flax-leaved Monardella. Fig. 4442. 



Monardella linoides A. Gray, Proc. Amer. Acad. 11: 101. 1876. 

 Monardella anemonoides Greene, Pittonia 5:86. 1902. 

 Madronella linoides Greene, Leaflets Bot. Obs. 1: 169. 1906. 



Stems several from a woody base, the branches erect, 25-45 cm. high, almost silvery with a 

 dense puberulence of microscopically minute retrorsely appressed hairs. Leaves linear to linear- 

 lanceolate, 1.5-2.5 cm. long, acute, narrowed at base to a short winged petiole or subsessile, 

 microscopically puberulent and silvery on both sides; bracts ovate to oblong-ovate, 10-15 mm. 

 long, membranous, white or rarely slightly tinged with light rose, prominently veined, puberulent 

 and obscurely scurfy, soft-ciliate on the margins ; calyx 8-9 mm. long, short-pubescent, the teeth 

 hirsute-pubescent ; corolla pale rose, 12-15 mm. long. 



Dry slopes. Upper Sonoran Zone; southern Sierra Nevada, desert ranges of Mono and Inyo Counties, and 



