756 SCROPHULARIACEAE 



51. Penstemon Stephensii Brandg. Stephens' Penstemon. Fig. 4688. 



Penstemon Stephensii Brandg. Zoe 5: 151. 1903. 



Penstemon Clevelandii var. Stephensii Munz & Jtn. Bull. Torrey Club 49: 41. 1922. 



Stems few to several, 3-10 dm. high. Leaves thinnish, mostly finely and sharply denticulate, 

 the several upper cauline pairs connate-perfoliate, forming disks up to 10 cm. long and 4 cm. 

 broad; thyrsus strict, subsecund, sparingly glandular, 1-3.5 dm. long, 3-4 cm. broad; calyx 

 3-4.5 mm. high, the lobes broadly ovate to subrotund, abruptly acute; corolla rose to pink- 

 lavender, without prominent guide lines, 17-22 mm. long, 4-6 mm. wide pressed, essentially 

 tubular or the throat slightly dilated, the tube about equaling the throat, the quadrate lobes 

 rotately spreading, the limb glandular-puberulent but not pilose within; anther-sacs peltately 

 explanate, glabrous, as broad as or broader than long; staminode included, glabrous. 



Rocky slopes, Lower Sonoran Zone; Kingston and Providence Mountains, eastern Mojave Desert, California. 

 Type locality: Providence Mountains. May-June. 



52. Penstemon incertus Brandg. Mojave Penstemon. Fig. 4689. 



Penstemon incertus Brandg. Bot. Gaz. 27:454. 1899. 



Penstemon fruticiformis var. incertus Munz & Jtn. Bull S. Calif. Acad. 23: 33. 1924. 



In habit similar to P. fruticiformis, usually shrubby, forming broad clumps in age, 6-8 dm. 

 high, the numerous stems branching below, the herbage glabrous and glaucous. Leaves narrowly 

 linear-lanceolate; thyrsus lax, moderately glandular; calyx 5-7 mm. high, the lobes lance-ovate 

 to nearly rotund ; corolla violet with a reddish cast or purple, the limb deep blue, without guide 

 lines, 25-28 mm. long, 8-11 mm. wide pressed, the rather long tube gradually expanding into 

 the ample throat, strongly 2-lipped, the lips reflexed, the lower one villous at base ; anther-sacs 

 divaricate, not explanate, minutely denticulate-ciliolate at suture ; staminode well included, short, 

 straight, densely bearded almost throughout. 



Dry sandy or rockv benches, Sonoran Zones; western borders of the Mojave Desert, California, from eastern 

 base of the Sierra Nevada and ArKUS Mountains, Inyo County, to Antelope Valley and base of the San Ber- 

 nardino Mountains. Type locality: Walker Pass. May-June. 



53. Penstemon spectabilis Thurb. Notable Penstemon. Fig. 4690. 



Penstemon spectabilis Thurb. ex A. Gray, Pacif. R. Rep. 4: 119. 1856. 



Stems several from the base, erect, 8-12 dm. high, the herbage green or glaucescent and 

 glabrous throughout. Leaves coarsely serrate, the lower broadly oblanceolate to ovate, 4-10 cm. 

 long, 2-5 cm. broad, the upper connate-perfoliate ; thyrsus lax, often half as tall as the plant ; 

 calyx 4-7 mm. high, the lobes lance-ovate to orbicular ; corolla lavender-purple with blue lobes, 

 whitish within, 25-33 mm. long, 8-12 mm. wide pressed, the tube rather abruptly expanding 

 into the ample throat, the limb strongly bilabiate, the upper lip nearly erect, the lower lip reflexed, 

 sometimes feebly bearded at base; anther-sacs not explanate, twice as long as wide, scabro- 

 ciliolate at suture ; staminode glabrous at tip. n = 8. 



Dry washes and hillsides. Upper Sonoran Zone; eastern Los Angeles County, California, to northern 

 Lower California. Type locality: San Pasqual, San Diegro County. April-June. 



Penstemon spectabilis subsp. subviscosus Keck, Amer. Midi. Nat. 18: 818. 1937. (Penstemon spectabilis 

 var. subviscosus McMinn, 111. Man. Calif. Shrubs 513. 1939.) Pedicels and calyces glandular-pubescent. The 

 common form in California from the Liebre and Santa Monica Mountains to the outwash fans of the San Gabriel 

 and San Bernardino Mountains. Type locality: Claremont, Los Angeles County. 



X Penstemon Parishii (P. centranthifolius X spectabilis) (A. Gray) Keck, Amer. Midi. Nat. 18: 818. 

 1937. (Penstemon Parishii A. Gray, Proc. Amer. Acad. 17:228. 1882; P. spectabilis var. Gilmanii Jepson, 

 Man. Fl. PI. Calif. 912. 1925.) Habit of P. centranthifolius; herbage glaucescent and glabrous throughout; 

 leaves entire to shallowly serrate, the uppermost clasping but distinct; thyrsus virgate; corolla red-purple, the 

 gradually ampliate throat 6-9 mm. wide. This hybrid is not uncommon in the region where the t\vo parents 

 occur together, from Los Angeles County to San Diego County, California. Type locality: "S.E. California, in 

 the Cucamonga Mountains." 



54. Penstemon centranthifolius Benth. Scarlet Bugler. Fig. 4691. 



Penstemon centranthifolius Benth. Scroph. Indicae 7. 1835. 



Glabrous and glaucous throughout ; stems several, virgate, 3-12 dm. high. Leaves entire, 

 the basal spatulate, petiolate, the cauline linear-lanceolate to ovate-lanceolate, the upper pairs 

 often auriculate-clasping. 4-10 cm. long, 1-3 cm. wide; thyrsus virgate, half as tall as the plant, 

 the peduncles erect ; calyx 3-6 mm. high, the lobes ovate to orbicular, abruptly acute, the margin 

 broadly scarious, entire to erose ; corolla scarlet, 25-33 mm. long, 4.5-6 mm. wide, tubular, the 

 lobes scarcely spreading, glabrous without and within ; anther-sacs peltately explanate ; stami- 

 node glabrous, n = 8. 



Dry slopes and outwash fans, Upper Sonoran Zone; Coast Ranges of California from Lake County to San 

 Diego County, south to the Sierra San Pedro Martir, Lower California. Type locality: "Nova California." 

 April-July. 



55. Penstemon utahensis Eastw. Utah Bugler. Fig. 4692. 



Penstemon utahensis Eastw. Zoe 4: 124. 1893. 

 Penstemon Eastwoodiae Heller, Muhlenbergia 1: 4. 1900. 



Stems several, glabrous, virgate, 3-6 dm. high. Leaves coriaceous, glaucous, glabrous, or 

 the basal scabridulous or scabrid-ciliolate, the basal lanceolate, tapering to the petiole, the cauline 

 lance-oblong, broadest at the clasping base, 3-8 cm. long, 5-15 mm. wide; thyrsus racemiform, 

 glabrous ; calyx 3-5 mm. high, the lobes ovate to orbicular, abruptly acute, the broadly scarious 



