742 SCROPHULARIACEAE 



explanate, 0.5-0.7 mm. long; staminode reaching orifice, lightly to rather densely tufted at 

 apex with short golden hairs, n = 24. 



On volcanic ash, in forest openings, Transition and Canadian Zones; Cascade Range, from Mount Adams, 

 Washington, to Three Sisters, Oregon. Type locality: Blue Grass Ridge, Mount Hood. June-Sept. 



12. Penstemon heterodoxus A. Gray. Sierran Penstemon. Fig. 4649. 



Penstemon heterodoxus A. Gray, Syn. Fl. N. Amer. 2i:269. 1878. 



Penstemon gentculatus Greene, Pittonia 3: 310. 1898. 



Penstemon alsinoides Greene, Leaflets Bot. Obs. 1 : 162. 1906. 



Penstemon depressus Greene, loc. cit. 



Penstemon procerus f. gentculatus Smiley, Univ. Calif. Pub. Bot. 9: 325. 1921. 



Penstemon confertus var. gentculatus Jepson, Man. Fl. PI. Calif. 914. 1925. 



Stems slender, mostly 8-15(-25) cm. high, the basal rosette developed. Leaves deep green, 

 thin, glabrous, the basal linear-oblanceolate to spatulate, 1-4 cm. long including the very slender 

 petiole as long as blade, 4-8 mm. vv^ide, the cauline oblanceolate to spatulate-oblong below and 

 narrowly to broadly lanceolate above, up to 3 cm. long and 8 mm. wide ; thyrsus usually reduced 

 to a subcapitate cluster, or of 2-4 distinct clusters, dense, very glandular ; calyx 3-6 mm. high, 

 the oblong lobes abruptly narrowed to the short acute tip; corolla deep blue-purple, 10-16 mm. 

 long, gradually ampliate, the palate prominently brownish-yellow-bearded; anther-sacs broadly 

 ovate, boat-shaped, rarely explanate, 0.7-1 mm. long; staminode included, bearded with short 

 stiff yellow hairs apically or sometimes glabrous, m = 8. 



Rocky slopes and alpine meadows, Hudsonian and Arctic-Alpine Zones; Sierra Nevada of California from 

 Plumas County to Mount Whitney; White Mountains; adjacent Nevada. Type locality: "High mountain near 

 Donner Pass, in the Sierra Nevada, California." July-Aug. 



Penstemon heterodoxus subsp. cephalophorus (Greene) Keck, Amer. Midi. Nat. 33: 165. 1945. {Penste- 

 mon ccphalophorus Greene, Leaflets Bot. Obs. 1: 79. 1904; P. glastifolius Greene, op. cit. 162. 1906.) Stems 

 stouter, mostly 15-40 cm. high; basal leaves 3-7 cm. long, 6-12 mm. wide, the middle cauline up to 6 cm. long 

 and 12 ram. wide. Southern Sierra Nevada in Fresno and Tulare Counties largely to the west of the Kern 

 River; at lower elevations than the typical form. Type locality: Summit Lake, Tulare County, California, 



13. Penstemon shastensis Keck. Shasta Penstemon. Fig. 4650. 



Penstemon shastensis Keck, Amer. Midi. Nat. 33: 165. 1945. 



Stems slender to rather stout, 2-5 dm. high, the basal rosette well developed. Leaves deep 

 green, rather thin, glabrous, the basal elliptic, obtuse, 3-6 cm. long including the long narrow 

 petiole, the lower cauline the largest, lance-oblong, up to 9 cm. long and 16 mm. wide; thyrsus 

 of 2-6 dense clusters or sometimes reduced to one, moderately glandular-pubescent; calyx 

 2.5-5 mm. high, the lance-oblong lobes not abruptly tipped, acuminate; corolla blue-purple, 

 10-13 mm. long, like heterodoxus; anther-sacs ovate, boat-shaped, 0.6-0.7 mm. long; staminode 

 reaching orifice, densely golden bearded at apex, n = 16. 



In meadows. Transition and Canadian Zones; California, Siskiyou and adjacent Modoc Counties to Shasta 

 County. Type locality: Grass Lake, Siskiyou County. June-July. This tetraploid species was formerly confused 

 with the diploid P. heterodoxus. 



14. Penstemon glaucinus Pennell. Blue-leaved Penstemon. Fig. 4651. 



Penstemon glaucinus Pennell, Notulae Naturae No. 71: 10. 1941. 



Stems slender, 3-3.5 dm. high, the herbage glaucous, the basal rosette well developed. Leaves 

 thickish, the basal elliptic to broadly spatulate, 2-6 cm. long including the slender petiole, 7-13 

 mm. wide, the cauline broadly oblong to lance-oblong ; thyrsus glandular-pubescent, of 2-4 more 

 or less remote clusters; calyx 3.5-6 mm. high, the lanceolate to ovate-oblong lobes attenuate, 

 the entire or erosulate margin prominently scarious ; corolla blue-purple, 12-15 mm. long, 

 gradually ampliate, the limb expanded or upper lip erect, the palate moderately bearded with 

 prominent yellow hairs; anther-sacs ovate, boat-shaped, 0.65-0.95 mm. long; staminode in- 

 cluded, densely bearded with short golden hairs or glabrous. 



Lodgepole pine forest, Canadian Zone; Campbell Lake, east of Gearhart Mountain, Lake County, Oregon 

 (the type locality), and south slope of Gearhart Mountain, both known collections made by E. I. Applegate. 

 July. 



15. Penstemon attenuatus Dougl. Taper-leaved Penstemon. Fig. 4652. 



Penstemon attenuatus Dougl. ex. Lindl. Bot. Reg. IS: pi. 1295. 1829. 



Penstemon propinquus Greene, Leaflets Bot. Obs. 1: 166. 1906. 



Penstemon confertus var. attenuatus M. E. Jones, Contr. West. Bot. No. 12: 62. 1908. 



Penstemon Netsonae Keck & Thomps. Rhodora 37 : 419. 1935. 



Stems rather slender, 3-6 (-9) dm. high, the basal rosette well developed. Leaves deep 

 green, usually entire or sometimes finely denticulate, glabrous, the basal linear-lanceolate to 

 oval, 4-10(-17) cm. long including the slender petiole, 7-20(-45) mm. wide, the cauline gradually 

 or abruptly reduced, the upper amplexicaul ; thyrsus glandular-pubescent, strict, of 3-7 clusters ; 

 calyx 4-7 mm. high, the lobes mostly lanceolate, with relatively narrow and entire scarious 

 margin ; corolla pale yellow or blue-purple to violet, mostly 14-20 mm. long, ampliate, rather 

 obviously bilabiate, the palate bearded with whitish hairs ; anther-sacs ovate, boat-shaped, 

 dehiscent through the connective and quite to the apex, mostly 0.8-1.2 mm. long; staminode 

 reaching orifice, bearded with relatively long golden hairs toward apex, n = 24. 



Openings in pine woods, Arid Transition Zone; central and eastern Washington (Chelan and Spokane 



