748 SCROPHULARIACEAE 



pale blue or lavender to nearly white. To the east of the species in the Arid Transition Zone; Snake River 

 watershed of eastern Oregon and adjacent Idaho. Type locality: "Mountains of E. Oregon, Cusick—. 



Penstemon Gairdneri subsp. hians (Piper) Keck, Amer. Midi. Nat. 23:597 1940 (Penstemoyt Gaird- 

 neri var hians Piper, Bull. Torrey Club 27:396. 1900.) Leaves alternate, short and narrow as in the 

 speciesrcorX rose^urple throughout, up to 25 mm. long, the limb usually 15-20 mm. across with widely 

 spreading lobes. Rocky prairies of central Washington from Chelan and Douglas Counties to Klickitat County. 

 Type locality: eastern Washington. 



29. Penstemon deustus Dougl. Hot-rock Penstemon. Fig. 4666. 



Penstemon deustus Dougl. ex. Lindl. Bot. Reg. 16: pi. 1318. 1830. 

 Penstemon deustus var. suffrutescens Henderson, Rhodora 33: 206. 1931. 

 Penstemon deustus var. Savagei Henderson, loc. cit. 



Stems woody and much branched below, forming clumps 2-6 dm. high, erect, glabrous or 

 glandular-puberulent. Leaves bright green, coarsely dentate-serrate, those of sterile shoots 

 1-5 cm. long, 6-20 mm. wide, short-petiolate, those of fertile shoots linear-lanceolate to elliptic- 

 ovate, sessile' or clasping, all acute to acuminate ; thyrsus strict, sparingly glandular ; calyx-lobes 

 lanceolate to ovate-attenuate; corolla ochroleucous, prominently marked with purplish guide 

 lines, 10-16 mm. long, nearly tubular, the upper lip shorter than the lower, sparingly glandular 

 without and within; anther-sacs orbicular, explanate, widely divaricate, 0.7 mm. long; staminode 

 reaching orifice, usually glabrous, sometimes short-bearded toward apex, n — 8. 



Dry rocky ground, mainly Arid Transition Zone; Columbia Basin, easterii Washington, to western Glenn 

 County and the central Sierra Nevada, California, east to Wyoming. Type locality: Native of North-west 

 America, where it was found by Mr. Douglas on scorched, rocky plains, in the interior. May-July. 



Penstemon deustus subsp. sudSns (M. E. Jones) Pennell & Keck, Amer. Midi. Nat. 23: 600. 1940. 

 (Penstemon sudans M. E. Jones, Contr. West. Bot. No. 8:37. 1898.) Herbage and corol a pr9minently 

 glandular-pubescent. Arid Transition Zone; common in volcanic soils in Lassen County, Calilornia. lype 

 locality: "between Amedee and Susanville." 



Penstemon deustus subsp. heterander (Torr. & Gray) Pennell & Keck, Amer. Midi. Nat. 23: 603. 1940. 

 (Penstemon heterander Torr. & Gray, Pacif. R. Rep. 2=: 123. 1855.) Very woody, with reduced stems; leaves 

 mostly narrow and finely toothed, glabrous, glaucescent; corolla obscurely viscid-puberulent without atid 

 glabrous within. Occupying the triangle from south central Oregon, to northeastern California, and north- 

 western Nevada to the exclusion of the typical form. Type locality: apparently in eastern Shasta County, 

 California. 



30. Penstemon variabilis Suksd. Variable Penstemon. Fig. 4667. 



Penstemon variabilis Suksd. Deutsch. Bot. Monatss. 18: 153. 1900. 

 Penstemon paniculatus Howell, Fl. N.W. Amer. 513. 1901. 



Closely resembling P. deustus in general habit and in the form and appearance of the 

 flower. Leaves ternate, quaternate, or opposite, or becoming somewhat scattered, narrowly 

 linear to lance-oblong, 2-8 mm. wide, quite entire to finely serrate toward the apex : inflorescence 

 usually branched, sometimes strict ; calyx-lobes glandular-pubescent to rarely glabrous ; corolla 

 10-1 2 (-15) mm. long, rather strongly glandular-pubescent; staminode usually bearded apically. 

 n=16. 



Open rocky slopes. Arid Transition Zone; Klickitat County, Washington, to Grant and Deschutes Coun- 

 ties, Oregon. Type locality: in a ravine east of the Klickitat River. June-July. 



31. Penstemon Tracy i Keck. Tracy's Penstemon. Fig. 4668. 



Penstemon Tracyi Keck, Amer. Midi. Nat. 23: 603. 1940. 



Suffrutescent subshrub 8-12 cm. high, the light green glaucescent herbage glabrous through- 

 out. Leaves coriaceous, cuneate-oblong or oval to orbicular, mostly entire, some finely denticu- 

 late, those of the basal rosette numerous, short-petiolate, the cauline usually tapering to a sessile 

 base; thyrsus contracted, dense, 2-4 cm. long, of 2 or 3 clusters; calyx 2.5-3 mm. high, 

 the lobes ovate, acute, with narrowly hyaline erosulate margin; corolla pink, 11-13 mm. long, 

 tubular, the palate densely villous, the limb small; anther-sacs orbicular, explanate, opposite, 

 0.4 mm. long; staminode included, sparsely bearded toward apex. 



A very distinctive species known only from the type locality, at the head of White's Creek, Devil's Canyon 

 Mountains, Trinity County, California, where it grows in rock crevices at 7,000 feet altitude. July-Aug. 



32. Penstemon albomarginatus M. E. Jones. White-margined Penstemon. 



Fig. 4669. 



Penstemon albomarginatus M. E. Jones, Contr. West. Bot. No. 12:61. 1908. 



Stems 15-30 cm. high, several from an elongated fleshy deeply buried root, the whole plant 

 pallid, glaucescent, glabrous. Leaves entire, spatulate to narrowly obovate, 2.5-6 cm. long 

 together with the calyx-lobes narrowly bordered with a white scarious more or less scabrid 

 margin; thyrsus leafy, 5-12 cm. long; calyx-lobes linear-lanceolate to broadly ovate-oblong; 

 corolla lavender-pink, whitish ventrally, with purple guide lines within, 13-18 mm. long, up to 

 5 mm. wide at throat, the two palatal ridges bearded with flattened yellow hairs, anther-sacs 

 broadly ovate, explanate; staminode reaching orifice, glabrous. 



Confined to deep sand. Lower Sonoran Zone; Mojave Desert of eastern California, southern Nevada, and 

 northwestern Arizona. Type locality: Goodsprings, Nevada. March-May. 



