800 SCROPHULARIACEAE 



5. Synthyris lanuginosa (Piper) Pennell & Thompson. Woolly Synthyris. 



Fig. 4797. 



Synthyris pinnatifida subsp. lanuginosa Piper, Contr. U.S. Nat. Herb. 11: 504. 1906. 

 Synthyris lanuginosa Pennell & Thompson, Proc. Acad. Phila. 85: 93. 1933. 



Lanuginous-canescent throughout, even upon the capsule, but the sepals glabrescent. Leaf- 

 blades bi- to tri-pinnatifid, the mid-portion linear, the primary segments about 4 pairs, each with 

 1 or 2 pairs of segments which are again toothed, or these sometimes more irregularly developed, 

 the ultimate segments acuminate and tending to be callose ; flowering stems exceeding the foliage, 

 scapose or with a few small bracts below inflorescence, the spiciform raceme 1-5 cm. long, with 

 pedicels 1-3 mm. long, much shorter than the ovate or oblanceolate bracts; sepals oblong- 

 lanceolate, 4 mm. long ; corolla 5-6 mm. wide, nearly rotate, the narrow entire lobes longer than 

 the tube ; filaments 4 mm. long ; style 4-5 mm. long ; capsule 4-5 mm. long, obcordately notched, 

 4 mm. wide ; seeds flat. 



Gravelly alpine summits, Arctic- Alpine Zone; Olympic Mountains of western Washington. Type locality: 

 Olympic Mountains, Washington. June. 



23. BESSEYA Rydb. Bull. Torrey Club 30: 279. 1903. 



Perennial glandless herbs, with radical cordate-ovate leaf-blades and erect flowering 

 stems that bear proximally several to many bracts on the spiciform inflorescence. Brac- 

 teoles none. Sepals 4 (in ours), united laterally near base. Corolla (in ours) lacking. 

 Stamens 2 (the upper pair). Stigmas united, minutely capitate. Capsule flattened, locu- 

 licidal. Seeds numerous, flattened. [Named in honor of Charles E. Bessey, American 

 botanist.] 



Species 9, of the western United States and the Great Lakes Region. Type species, Synthyris alpina A. Gray. 



1. Besseya rubra (Dougl.) Rydb. Red Besseya. Fig. 4798. 



Gytnnandra rubra Dougl. ex Hook. Fl. Bor. Amer. 2; 103. 1838. 

 Synthyris rubra Benth. in A. DC. Prod. 10: 455. 1846. 

 Wulfenia rubra Greene, Erythea 2: 83. 1894. 

 Besseya rubra Rydb. Bull. Torrey Club 30: 280. 1903. 

 Lunellia rubra Nieuwl. Amer. Midi. Nat. 3: 189. 1914. 



Plant loosely hairy, but the leaves becoming glabrescent, the inflorescence villulose in anthesis 

 with the capsule short-pubescent and in age glabrescent. Leaf-blades crenate-dentate, paler 

 beneath, truncate or slightly cordate at base ; flowering stems exceeding the foliage, proximally 

 with about 10 small bract-leaves, the distal spike-like raceme becoming 10-20 cm. long, its 

 lowest pedicels 2-3 mm. but most pedicels less than 1 mm. long; sepals 3-4 mm. long, oblong, 

 rounded ; filaments 4-5 mm. long, dark red ; style 4-5 mm. long ; capsule 5-6 mm. long, rounded, 

 6-7 mm. wide. 



Sandy or rocky prairie or open woodland. Arid Transition Zone; eastern Washington and eastern Oregon to 

 western Montana and southwestern Idaho. Type locality: northwestern Montana. April-May. 



24. PARENTUCELLIA Viviani, Fl. Libyc. Spec. 31. 1824. 



Erect annual or biennial hairy herbs, with opposite leaves and a spike-like raceme of 

 flowers. Bracteoles none. Calyx wath 4 lanceolate lobes. Corolla 2-lipped, its upper lip 

 galeate, with lobes united to apex, its lower lip with 2-ridged palate and with short spread- 

 ing lobes. Stamens 4, didynamous, the anthers lanose, with cells equivalent and mucronate- 

 tipped. Capsule cylindric, acute, loculicidal, the upper cell slightly the larger. Seeds el- 

 lipsoid-oblong, smooth. [Named in honor of Tomaso Parentucelli, founder of the botanic 

 garden at Rome.] 



Two species, of the Mediterranean Region, the following naturalized in western North America and in 

 southern South America. Type species, Parentucellia floribunda Viviani, of Libya. 



1. Parentucellia viscosa (L.) Caruel. Yellow Parentucellia. Fig. 4799. 



Bartsia viscosa L. Sp. PI. 602. 1753. 



Parentucellia viscosa Caruel in Pari. Fl. Ital. 6: 482. 1885. 



Plant glandular-pubescent throughout (except between ribs on lower leaf-surface), the stem 

 3-5 dm. tall. Leaf-blades saliently dentate, rounded to sessile bases ; pedicels less than 2 mm. 

 long; corolla 16-17 mm. long, yellow; capsule 8 mm. long, distally brown-hirsute; seeds 

 0.3 mm. long. 



Moist waysides, near the coast, Oregon and northern California. April-July. 



25. BELLArDIA All. Fl. Ped. 1: 61. 1785. 



Erect annual hairy herbs, with opposite leaves and a spike-like raceme of purple 

 flowers. Bracteoles none. Calyx with 4 short lobes. Corolla 2-lipped, its upper lip galeate, 

 with lobes united to apex, its lower lip with 2-ridged palate and with short spreading 



