ON CARYOKINESIS. 1/9 



the intra-miclear archoplasmic filaments {E) ; while those 

 lying outside of the elliptical body are the extra-nuclear 

 arcJwplasmic filaments {B). 



The elliptical portion of the figure consists of three 

 parts, two terminal and one intermediate. The terminal 

 portion, which presents different optical properties from 

 the intermediate part, consists of a hemispherical mass 

 of a slightly stainable, semi-liquid substance, which I 

 believe to be the nuclear-sap of the original nucleus. 

 Into this part the archoplasmic rays extend, as has 

 already been mentioned. The two terminal masses of 

 stainable substance are separated from the intermediate 

 non-stainable bundle of filaments by parallel chromatic 

 '* plates " (;/), (;/), — the chromosomes (Waldeyer) of 

 the original nucleus. The non-stainable intermediate 

 filaments above referred to are the interzonal archoplas- 

 mic filaments {D), — ''interzonal filaments" of Mark, 

 "filaments reunissants " of van Beneden, " gubernacu- 

 lum"of Maupas, " Verbindungschlauch" of Strasburger, 

 ''connective filaments," " Verbindungsfaden," etc., of 

 authors. 



One plate of chromosomes goes to one daughter 

 nucleus, and the other to another. The cytoplasm 

 accumulates around each, and there follows a separation 

 into two cells, each with its distinct nucleus. 



If one examine a nucleus at a tolerably early stage of 

 caryokinesis, one will see a phenomenon such as is 

 shown in Fig. 3. The nucleus with a network of chromo- 

 somes is intercepted between two archoplasmic spheres. 

 More than this, however. That portion of the archo- 

 plasmic rays which falls on the surface of the nucleus 

 presses that part inward and so flattens that side of the 



